Such organic combination, as opposed to mere commixture, is altogether
hypothetical, but it may be worth while to glance at some of its
implications. If it be analogous to chemical combination, the products
would be of a definite nature; in other words, the variations would be
in definite directions. Selection and elimination would not have to deal
with variations in any and all directions, but would have presented to
them variations specially directed along certain lines determined by the
laws of organic combination. As Professor Huxley has said, "It is quite
conceivable that every species tends to produce varieties of a limited
number and kind, and that the effect of natural selection is to favour
the development of some of these, while it opposes the development of
others along their predetermined line of modification." Mr. Gulick[BQ]
and others have been led to believe in a tendency to divergent evolution
residing in organic life-forms. Such a tendency might be due to special
modes of organic combination giving rise to particular lines of
divergence. Again, we have seen that some naturalists believe that
specific characters are not always of utilitarian significance. But, as
was before pointed out, on the hypothesis of all-round variation, there
is nothing to give these non-useful specific characters fixity and
stability, nothing to prevent their being swamped by intercrossing. If,
however, on the hypothesis of combination, we have definite organic
compounds, instead of, or as well as, mere hereditary mixtures; if, in
other words, variations take definite lines determined by the laws of
organic combination (as the nature and properties of chemical compounds
are determined by the laws of chemical combination), then this
difficulty disappears. There is no reason why a neutral divergence--one
neither useful nor deleterious--should be selected or eliminated. And if
its direction is predetermined, there is no reason why it should not
persist, though, of course, it will not be kept at a high standard by
elimination. It has again and again been pointed out as a difficulty in
the path of natural selection that, in their first inception, certain
characters or structures cannot yet be of sufficient utility to give the
possessor much advantage in the struggle for existence. If, however,
these be definite products of organic combination, this difficulty also
disappears. So long as they are not harmful, they will not be
eliminated, and by fortunate combinations will progress slowly until
natural selection gets a hold on them and pushes them forward,
developing to the full the inherent tendency. Finally, we must notice
that, on this hypothesis, our conception of panmixia, or intercrossing,
would have to be modified. As generally held, this doctrine is based
upon hereditary mixture, not organic combination. It is a doctrine of
means and averages. There is a good deal of evidence that intercrossing
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