does not, at least in all cases, produce mean or average results. And
according to the hypothesis of organic combination, it need not always
do so. According to this hypothesis, then, divergent modifications might
arise and be perpetuated without the necessity of isolation. Sterility
might result from the fact that divergence had been carried so far that
organic combination was no longer possible; reversion, due to
intercrossing, from the fact that combinations long rendered impossible
by the isolation of the necessary factors in distinct varieties, are
again rendered possible when these varieties interbreed.
On this hypothesis of organic combination, to which we shall recur in
the chapter on "Organic Evolution," the varied forms of animal life are
the outcome of definite organic products with definite organic
structure, analogous to the definite chemical compounds with definite
crystalline and molecular structure; and the analogy between the
regeneration of hydra and the reconstruction of a crystal is carried on
a step further. I do not say that I am myself at present prepared to
adopt the hypothesis, at least in this crude form; but it is, perhaps,
worth a passing consideration. Its connection with Mr. Herbert Spencer's
doctrine of physiological units is obvious. The analogy there is with
crystallization; here it is with chemical combination.
We must now return to the point which gave rise to this digression, and
repeat that mere hereditary commixture in the union of ovum and sperm
cannot give rise to new characters or raise existing structures (1)
where there is free intercrossing beyond the mean of the species, and
(2) where there is rigorous elimination beyond the existing maximum of
the species. Variations beyond this existing maximum must be due to some
other cause.
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