Nor can it be questioned that variations in the time occupied in
reaching maturity would, other things equal, not a little affect the
chances of success of an organism in the competition of life. Hence we
have the phenomena of what may be termed acceleration and retardation in
development. These terms have, however, been used by American
zoologists, notably Professors Hyatt and Cope, in a somewhat different
and wider sense; for they include not merely time-changes, but also the
loss of old characters or the acquisition of new characters. "It is
evident," says Professor Cope, "that the animal which adds something to
its structure which its parents did not possess has grown more than
they; while that which does not attain to all the characteristics of its
ancestors has grown less than they." "If the embryonic form be the
parent, the advanced descendant is produced by an increased rate of
growth, which phenomenon is called 'acceleration'; but if the embryonic
type be the offspring, then its failure to attain the condition of the
parent is due to the supervention of a slower rate of growth; to this
phenomenon the term 'retardation' is applied." "I believe that this is
the simplest mode of stating and explaining the law of variation: that
some forms acquire something which their parents did not possess; and
that those which acquire something additional have to pass through more
numerous stages than their ancestors; and those which lose something
pass through fewer stages than their ancestors; and these processes are
expressed by the terms 'acceleration' and 'retardation.'"[DO]
It is clear, however, that we have here something more than acceleration
and retardation of development in the ordinary sense of these words. It
would be, therefore, more convenient to use the term "acceleration" for
the condensation of _the same series_ of developmental changes into a
shorter period of time; "retardation" for the lengthening of the period
in which _the same series_ of changes are effected; and "arrested
development" for those cases in which the young are born in an immature
or embryonic condition. Whether there is any distinct tendency, worthy
of formulation as a law, for organisms to acquire, as a result of
protracted embryonic development, definite characteristics which their
ancestors did not possess, I think very questionable. If so, this will
fall under the head of the origin of variations.
Public-domain text, read in full here on John Shaqi.
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