That acceleration, in the sense in which I have used the term, does
occur as a variation is well known. "With our highly improved breeds of
all kinds," says Darwin,[DP] "the periods of maturity and reproduction
have advanced with respect to the age of the animal; and in
correspondence with this, the teeth are now developed earlier than
formerly, so that, to the surprise of agriculturalists, the ancient
rules for judging of the age of an animal by the state of its teeth are
no longer trustworthy." "Disease is apt to come on earlier in the child
than in the parent; the exceptions in the other direction being very
much rarer."[DQ] Professor Weismann contends that the time of
reproduction has been accelerated through natural selection, since the
shorter the time before reproduction, the less the number of possible
accidents. We may, perhaps, see in the curious cases of reproduction
during an otherwise immature condition, extreme instances of
acceleration. The axolotl habitually reproduces in the gilled, or
immature condition. Some species of insects reproduce before they
complete their metamorphoses. And the females of certain beetles
(_Phengodini_) are described by Professor Riley as larviform.[DR]
Precocity is variation in the direction of acceleration, and that
condensed development which is familiar in the embryos of so many of the
higher animals may be regarded as the result of variations constantly
tending in the same direction. That there are fewer examples of
retardation is probably due to the fact that nature has constantly
favoured those that can do the same work equally well in a shorter time
than their neighbours. But there can be no doubt that, accompanying that
fosterage and protection which is of such marked import in the higher
animals, there is also much retardation. And as bearing upon the
supposed law of variation as formulated by Messrs. Hyatt and Cope, it
should be noted that this retardation or _decreased_ rate of growth
leads to the production of the more advanced descendant.
_The Inheritance of Variations._
Public-domain text, read in full here on John Shaqi.
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