Although it is probably not functional in any existing form, mention
must here be made of the median or pineal eye. On the head of the common
slow-worm, or blind-worm, there is a dark patch surrounding a brighter
spot. This is the remnant of a median eye. It has been found in varying
states of degeneration in many reptiles (Fig. 34), and in a yet more
vestigial form in some fishes and amphibia. It is connected with a
curious structure, associated with the brain of all vertebrates, and
called the pineal gland. Descartes thought that this was the seat of the
soul; but modern investigation shows it to be a structure which has
resulted from the degeneration of that part of the brain which was
connected with the median eye. There is some reason to suppose that, in
ancient life-forms, like the Ichthyosaurus, and Plesiosaurus, and the
Labyrinthodont amphibians, it was large and functional. In any case,
there is a large hole in the skull (Fig. 35) through which the nervous
connection with the brain may have been established. The structure of
the eye is not similar to that of the lateral eye, but more like that of
some of the invertebrates.
To these invertebrates we must now turn.
[Illustration: Fig. 35.--Skull of _Melanerpeton_.
A Labyrinthodont amphibian from the Permian of Bohemia (after Fritsch).
× 4. Pa., the parietal foramen.]
* * * * *
Insects have eyes of two kinds. If we examine with a lens the head of a
bee, we shall see, on either side, the large compound or facetted eye;
but in addition to these there is on the forehead or vertex a triangle
of three small, bright, simple eyes, or ocelli. These ocelli, or
eyelets, differ, in different insects, as to the details of their
structure; but in general they consist of a lens produced by the
thickening of the integumentary layer which is at the same time rendered
transparent. Behind this lies the so-called vitreous body, composed of
transparent cells, and then follows the retina, in which there are a
number of rods, the recipient ends of which are turned towards the rays
of light, and not away from them as in the vertebrate. Spiders have from
six to eight ocelli, arranged in a pattern on the top of the head.
Facetted eyes are not found in them.
[Illustration: Fig. 36.--Eyes and eyelets of bee.
A. Drone. B. Worker.]
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