These facetted eyes, which are found in both insects and crustacea, have
apparently a more complex structure than the ocelli. Externally--in the
bee, for example--the surface is seen to be divided up into a great
number of hexagonal areas, each of which is called a facet, and forms
(in some insects, but not in all) a little lens. Of these the queen bee
has on each side nearly five thousand; the worker some six thousand; and
the drone upwards of twelve thousand; while a dragon-fly (_Æschna_) is
stated to have twenty thousand. Beneath each facet (in transverse
section, Fig. 37) is a crystalline cone, its base applied to the lens,
its apex embraced by a group of elongated cells, in the midst of which
is a nerve-rod which is stated to be in direct connection with the
fibres of the optic nerve. Dark pigment is developed around the
crystalline cones. And retinal purple is said to be present in the cells
which underlie it.
With regard to these facetted eyes there has been much discussion. The
question is--Is each facetted organ an eye, or is it an aggregate of
eyes? To this question the older naturalists answered confidently--An
aggregate. A simple experiment seems to warrant this conclusion. If the
facetted surface be cleared of its internal structures (the crystalline
cones, etc.) and placed under the microscope, each lens may, at a
suitable distance of the object-glass, be made to give a separate image
of such an object as a candle reflected in the mirror of the microscope.
If each lens thus gives an image, is not each the focussing apparatus of
a single eye? But a somewhat more difficult experiment points in another
direction. If the facetted cornea be removed _with the crystalline cones
still attached_ (Grenacher was able to do it with a moth's eye), and
placed under the microscope, when the instrument is focussed at the
point of the cone (where the nerve-rod comes), a spot of light, and not
an image, is seen. No image can be seen unless the microscope be
focussed for the centre of the cone; and here there are no structures
capable of receiving it and transmitting corresponding waves of change
to the "brain."
[Illustration: Fig. 37.--Eye of fly.
Transverse section through head. (After Hickson.)]
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