Now, if we study the process of respiration and that of circulation,
with which it is so closely associated, in other forms of life, we shall
find many differences in detail. In the bird, for example, the mechanism
of respiration is different. There is no diaphragm, and the lungs are
scarcely distensible. There are, however, large air-sacs in the abdomen,
in the thoracic region, in the fork of the merry-thought, and elsewhere.
These are distensible, and to reach them the air has to pass through the
lungs, and as it thus passes through the delicate tubes of the lungs, it
supplies the blood with oxygen and takes away carbonic acid gas. In the
frog there is no diaphragm, and there are no ribs. The lungs are hollow
sacs with honey-combed sides, and they are inflated from the mouth,
which is used as a force-pump for this purpose. In the fish there are no
lungs, respiration being effected by means of gills. In these organs the
blood is separated from the water which passes over them (being gulped
in by the mouth and forced out between the gill-covers) by only a thin
organic film, so that it can take up the oxygen dissolved in the water,
and give up to the water the carbonic acid it contains. In fishes, too,
we have only one receiver and one force-pump, the blood passing through
the gills on its way to the various parts of the body. In the lobster,
again, there are gills, but the mechanism by which the water is drawn
over them is quite different, and the blood passes through them on its
way to the heart, after passing through the various organs of the body,
not on its way from the heart, as in vertebrate fishes. The blood, too,
has no red blood-discs. In the air-breathing insects the mechanism is,
again, altogether different. The air, which obtains access to the body
by spiracles in the sides (see Fig. 1, p. 3), is distributed by delicate
and beautiful tubes to all parts of the organs; so that the oxygen is
supplied to the tissues directly, and not through the intervention of a
blood-stream. In the earthworm, on the other hand, there is a
distributing blood-stream, but there is no mechanism for introducing the
air within the body; while in some of the lowliest forms of life there
is neither any introduction of air within the body nor any distribution
by means of a circulating fluid. Beginning, therefore, with the surface
of the body simply absorbent of oxygen, we have the concentration of the
absorbent parts in special regions, and an increase in the absorbent
surface, either (1) by the pushing out of processes into the surrounding
medium, as in gills; or (2) by the formation of internal cavities,
tubes, or branching passages, as in lungs and the tracheal air-system of
insects.
What, then, is the essential nature of the respiratory process thus so
differently manifested? Clearly the supply of oxygen to the cellular
tissue-elements, and, generally closely associated with this, the
getting rid of carbonic acid gas.
Public-domain text, read in full here on John Shaqi.
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