This view of development is, however, the outcome of comparatively
modern investigation and perfected microscopical appliances. The older
view was that development in all cases is nothing more than differential
growth, that there is no differentiation of primitively similar into
ultimately different parts. Within the fertilized ovum of the horse or
bird lay, it was supposed, in all perfection of structure, a miniature
racer or chick, the parts all there, but too minute to be visible. All
that was required was that each part should grow in due proportion.
Those who held this view, however, divided into two schools. The one
believed that the miniature organism was contained within the ovum, the
function of the sperm being merely to stimulate its subsequent
developmental growth. The other held that the sperm was the miniature
organism, the ovum merely affording the food-material necessary for its
developmental growth. In either case, this unfolding of the invisible
organic bud was the _evolution_ of the older writers on organic life.
More than this. As Messrs. Geddes and Thomson remind us,[H] "the germ
was more than a marvellous bud-like miniature of the adult. It
necessarily included, in its turn, the next generation, and this the
next--in short, all future generations. Germ within germ, in ever
smaller miniature, after the fashion of an infinite juggler's box, was
the corollary logically appended to this theory of preformation and
unfolding."
Modern embryology has completely negatived any such view as that of
preformation, and as completely established that the evolution is not
the unfolding of a miniature germ, but the growth and differentiation of
primitively similar cell-elements. In different animals, as might be
expected, the manner and course of development are different. We may
here illustrate it by a very generalized and so to speak diagrammatic
description of the development of a primitive vertebrate.
[Illustration: Fig. 11.--Diagram of development. See text. The fine line
across G. indicates the plane of section shown in H.]
The ovum before fertilization is a simple spherical cell, without any
large amount of nutritive material in the form of food-yolk (A.). It
contains a nucleus. Previous to fertilization, however, in many forms of
life, portions of the nucleus, amounting to three parts of its mass, are
got rid of in little "polar cells" budded off from the ovum. The import
of this process we shall have to consider in connection with the subject
of heredity. The sperm is also a nucleated cell; and on its entrance
into the ovum there are for a short time two nuclei--the female nucleus
proper to the ovum, and the male nucleus introduced by the sperm. These
two unite and fuse to form a joint nucleus. Thus the fertilized ovum
starts with a perfect blending of the nuclear elements from two cells
produced by different parents.
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