Another mode of segregation arises also out of geographical conditions.
If variations of habits occur (and structure is closely correlated with
habit) such that certain individuals take to the mountains, others to
the plains or valleys; or that certain individuals take to the forests,
others to the open country; the probabilities are that the forest forms
will interbreed frequently with each other, but seldom with those in the
open, and so with the other varieties. The conditions of forest life or
mountain life being thus similar throughout a large area, and life being
through elimination slowly but surely adapted to its environment, there
might thus arise two distinct varieties scattered throughout the length
and breadth of the area, the one inhabiting the mountains, the other the
forests. In illustration of this mode of segregation, we may take the
case of two species of rats which have recently been found by Mr. C. M.
Woodford on one of the Solomon Islands. These two quite distinct species
are regarded by Mr. Oldfield Thomas as slightly modified descendants of
one parent species, the modifications resulting from the fact that of
this original species some individuals have adopted a terrestrial,
others an arboreal life, and their respective descendants have been
modified accordingly. Thus _Mus rex_ lives in trees, has broad
foot-pads, and a long rasp-like, probably semi-prehensile, tail; while
_Mus imperator_ lives on the ground, has smaller pads, and a short,
smooth tail. The segregation of these two species has probably been
effected by the difference of their mode of life, and each has been
adapted to its special environment through the elimination of those
individuals which were not in harmony with the condition of their life.
It is probable that this mode of segregation has been an important one.
And it is clear that in many cases competition would be a co-operating
factor in this process, weaker organisms being forced into otherwise
uncongenial habitats through the stress of competitive elimination, the
weaker forms not perishing, but being eliminated from more favoured
areas.
Protective coloration may also be a means of segregation. A species of
insects having no protective resemblance might vary in two
directions--in the direction of green tints, assimilating their hue to
that of vegetation; and in the direction of sandy or dull earthy
colours, assimilating them to the colour of the soil. In the one variety
elimination would weed out all but the green forms, and these would be
left to intercross. In the other variety, green forms would be
eliminated, dull-brown forms being left to interbreed. Stragglers from
one group into the other would stand a chance of elimination before
interbreeding was effected.[AO]
Public-domain text, read in full here on John Shaqi.
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