In the case of birds whose freedom of flight gives them a wide range,
sometimes almost a world-wide range, it would seem at first sight that
their facilities for interbreeding and intercrossing are so great that
divergence is well-nigh impossible. And yet the examples of divergence I
cited from Mr. Wallace were taken from birds, and it is well known that
divergence is particularly well shown in this class. But when the habits
of birds are studied attentively, it is found that, wide as is their
range, their breeding area is often markedly restricted. The sanderling
and knot range freely during the winter throughout the Northern
hemisphere; but their breeding area is restricted to the north polar
region. The interbreeding within this area keeps the species one and
homogeneous, notwithstanding its wide range, and, at the same time,
prevents intercrossing with allied species with different
breeding-grounds.
Another most important mode of segregation among animals arises out of
habitual or instinctive preferences. Where varieties are formed there is
a tendency for like to breed with like. In the Falkland Islands the
differently coloured herds of cattle, all descended from the same stock,
keep separate, and interbreed with each other, but not with individuals
outside their own colour-caste. If two flocks of merino sheep and heath
sheep be mixed together, they do not interbreed. In the Forest of Dean
and in the New Forest, the dark and pale coloured herds of fallow deer
have never been known to intermingle.[AP] Here we have a case of
selective _segregation through preferential mating_, and may find
therein the basis of sexual selection in its higher ranges as advocated
by Darwin.
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