But when we take a more extended view of the matter, and include secular
changes of climate, the possible range of variation in the struggle for
existence is seen to be enormously increased. It is well known to those
who have followed the progress of geology, that in early Kainozoic times
a mild climate extended to within the Arctic circle, while during the
glacial epoch much of the north temperate zone was fast locked in ice,
and the climate of the northern hemisphere was profoundly modified. The
animals in the north temperate zone were driven southwards.[AZ] Not only
was there much elimination from the severe climatic conditions, but the
migrants were driven southwards into areas already well stocked with
life, and the competition for means of subsistence in these areas must
have been rendered extremely severe. Elimination was at a maximum. Then
followed the withdrawal of glacial conditions. The increasing geniality
of the climate allowed an expansion of life within a given area, and the
withdrawal of snow and ice further and further north set free new areas
into which this expanding life could migrate and find subsistence. The
hard times of the glacial period were succeeded by good times of
returning warmth and an expanding area; and if, as some geologists
believe, there was an inter-glacial period (or more than one such
period) in the midst of the Great Ice Age, then hard times and good
times alternated during the glacial epoch.
Expansion and contraction of life-areas have also been effected again
and again in the course of geological history by elevations and
subsidences of the land. At the beginning of Mesozoic times much of
Europe was dry land. In Triassic and Rhætic times there were lakes in
England and in Germany, and a warm Mediterranean Sea to the south.
Subsidence of the European area brought with it a lessened land-area and
an increased sea-area: bad times and increased competition for land
animals; good times and a widening life-area for marine forms of life.
This continued, with minor variations, till its culmination in the
Cretaceous period. Then came the converse process: the land-areas
increased, the sea was driven back. A good time had come for terrestrial
life; the marine inhabitants of estuaries and inland seas felt the
pressure of increased competition in a lessening area. And so there
emerged the continental Europe of the beginning of the Kainozoic era.
And it is scarcely necessary to remind those who are in any degree
conversant with geology that during tertiary times there have been
alternate expansions and contractions of life-areas, marine and
terrestrial, the former bringing good times, the latter hard times and a
heightened struggle for existence.
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