Now, what would be the result of this alternation of good times and hard
times? During good times varieties, which would be otherwise unable to
hold their own, might arise and have time to establish themselves. In an
expanding area migration would take place, local segregation in the
colonial areas would be rendered possible, differential elimination in
the different migration-areas would produce divergence. There would be
diminished elimination of neutral variations, thus affording
opportunities for experimental combinations. In general, good times
would favour variation and divergence.
Intermediate between good times and hard times would come, in logical
order, the times in which there is neither an expansion nor a
contraction of the life-area. One may suppose that these are times of
relatively little change. There is neither the divergence rendered
possible by the expansion of life-area, nor the heightened elimination
enforced by the contraction of life-area.[BA] Elimination is steadily in
progress, for the law of increase must still hold good. Divergence is
still taking place, for the law of variation still obtains. But neither
is at its maximum. These are the good old-fashioned times of slow and
steady conservative progress. They are, perhaps, well exemplified by the
fauna of the Carboniferous period, and it is not at all improbable that
we are ourselves living in such a quiet, conservative period.
On the other hand, hard times would mean increased elimination. During
the exhibitions at South Kensington there were good times for rats. But
when the show was over, there followed times that were cruelly hard. The
keenest competition for the scanty food arose, and the poor animals were
forced to prey upon each other. "Their cravings for food," we read in
_Nature_, "culminated in a fierce onslaught on one another, which was
evidenced by the piteous cries of those being devoured. The method of
seizing their victims was to suddenly make a raid upon one weaker or
smaller than themselves, and, after overpowering it by numbers, to tear
it in pieces." Elimination by competition, passing in this way into
elimination by battle, would, during hard times, be increased. None but
the best organized and best adapted could hope to escape. There would be
no room for neutral variations, which, in the keenness of the struggle,
would be relatively disadvantageous. Slightly divergent varieties,
before kept apart through local segregation, would be brought into
competition. The weakest would in some cases be eliminated. In other
cases, the best-adapted individuals of each variety might survive. If
their experiments in intercrossing, should such occur, gave rise to
fertile offspring, more vigorous and better adapted than either
parent-race, these would survive, and the parent-forms would be
eliminated. But if such experiments in intercrossing gave rise to
infertile, weakly offspring, these would be eliminated. Thus sterility
between species would become fixed.
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