Wherever, during the preceding good
times, divergence had taken place in two different directions of
adaptation, and some intermediate forms, fairly good in both directions,
had been able to escape elimination, the chances are that these
intermediates would be in hard times eliminated, and the divergent forms
left in possession of the field. Wherever, during good times, a species
had acquired or retained a habit of flexibility, that habit would stand
it in good stead in the midst of the changes wrought by hard times; but
when it had, on the other hand, acquired rigidity (like the proverbially
"inflexible goose"), it would be at a disadvantage in the stress of a
heightened elimination.
The alternation of good times and hard times may be illustrated by an
example taken from human life. The introduction of ostrich-farming in
South Africa brought good times to farmers. Whereupon there followed
divergence in two directions. Some devoted increased profits to
improvements upon their farms, to irrigation works which could not
before be afforded, and so forth. For others increased income meant
increased expenditure and an easier, if not more luxurious, mode of
life. Then came hard times. Others, in Africa and elsewhere, learnt the
secret of ostrich-farming. Competition brought down profits, and
elimination set in--of which variety need hardly be stated.
I believe that the alternation of good times and hard times, during
secular changes of climate and alternate expansions and contractions of
life-areas through geological upheavals and depression of the land, has
been a factor of the very greatest importance in the evolution of varied
and divergent forms of life, and in the elimination of intermediate
forms between adaptive variations. It now only remains in this chapter
to say a few words concerning convergence, adaptation, and progress.
Public-domain text, read in full here on John Shaqi.
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