K. E. von Bär does not agree with Huxley in considering that there is
less difference between man and the gorilla than that which exists
between different species of apes. “There are,” Von Bär remarks,
“differences of various kinds among apes. In some the thumb is only
a stump; in others, as in the orang-utan, the fingers of the hinder
extremities are so long and curved that they cannot be extended on flat
ground; in many of the smaller apes this member is still more like a
hand than in the larger species, and the fingers can be easily spread
out on the ground. In this case the foot is of a much blunter form, and
is more flexible, so that the sole, which is properly turned inward,
can lie flat on the ground. The heavier the body of the animal, the
more sharply cut the structure of the foot must be, so that it does
not admit of the free movements which are possible in the hand. But
all these are only modifications of a climbing foot, or prehensile
member--that is, of a hand, not modifications of a foot resting firmly
on the ground and supporting the whole weight of the body.
“It must not be forgotten that the structure of the skeleton is subject
to mechanical laws, which may be traced through the whole series of
the animal world. This is readily apparent when we turn to the human
structure.
“The human foot rests for the greater part of its length on the
ground, that is to say, with the heel and centre of the foot, which
form together a firm arch. The tarsus consists of the astragalus, and
also of the _os calcis_, which in man form a very prominent part,
taking a backward and downward direction, and of five other bones.
The metatarsus consists of five bones, on which the five toes are
inserted. In man these metatarsal bones are considerably longer than
the separate phalanges. Thus, the arch on which man is supported in
an erect position extends from the heel to the extremities of the
metatarsal bones. The several bones are slightly movable, but they are
so firmly connected that they can diverge but little from each other,
unless muscular power is exerted. In order to press the toes upon the
ground, it is again necessary to exert the muscles. The arched instep
has this advantage, that the foot can take a better hold of the slight
inequalities of the ground. In a profile view of the skeleton of a
human foot, the shortness of the toes, in comparison with the length of
the arched instep, is very apparent. In any natural position, even when
man is not walking or standing, the sole of the foot is not turned
inwards, but downwards.... The toes of the gorilla take the form of
a hand, since the great toe stands separate like a thumb, while the
other toes are turned outwards. In the gorilla the tarsus is short,
and the heel is bent inwards. The several bones of the human foot are
undoubtedly present in the hind hand of a gorilla, but the organ is
changed into a prehensile organ or hand. The conditions are the same as
Public-domain text, read in full here on John Shaqi.
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