When the gorilla is provoked, he displays both rows of teeth, and opens
his mouth to utter sounds of fury, while making ready to fight. It is
well known that anthropoids are able to pout and project their lips;
and Darwin says that they do this, not only when they are slightly
teased, and are sullen or disappointed, but also when anything occurs
to make them uneasy.
I have often observed in chimpanzees a slight wrinkling of the region
of the nasal cartilage, and even a vibration in a lateral and upward
direction. In any case, the muscles which we have described as acting
on the nose and upper lip are exercised.
The platysma myoides, which extends in man from the lower row of teeth
to just below the clavicle, occupies about the same area in the gibbon
and in other apes (Fig. 50). In the chimpanzee, however, this muscle
extends as high as the zygomatic arch, or even higher. In the gorilla
also I observed that this part extends comparatively high on the face.
In chimpanzees, orangs, and gibbons the upper fibres of this muscle
seem to form the risorius. In one case the platysma myoides sent forth
a fasciculus, about 18 mm. in width, to the beginning of the lower
temporal ridges. In the gorilla I saw that the uppermost fibres of the
platysma myoides were partly covered by the risorius (Fig. 50, ~10~).
From the corresponding muscle in the orang the lower fibres tend far
backward, and are in connection with the deltoid muscle covering a
segment of the capsular ligament. This muscle wrinkles the skin of
the neck, and helps to draw down the lower jaw. In cases in which it
extends far in an upward direction, as in those we have cited, it
affects the lateral extension of the middle and lower skin on the faces
of these animals, as well as the grinning contortion of the corner of
the mouth. It may also have to do with the grumbling sound issuing from
the throat-pouch, which is uttered by the animal when agitated, as he
rapidly opens and closes his mouth.
The strong sterno-cleido-mastoid muscle found in these animals, and
especially in the orang and gibbon, can be divided without difficulty
into a sternal and clavicular portion. The two portions diverge from
each other in a downward direction. As Bischoff justly states, a
muscle not hitherto observed in man may be traced in all four species
of anthropoids, a muscle which extends from the external part of the
clavicle to the transverse process of the first cervical vertebra.
Bischoff has called it the musculus omocervicalis. It is found in other
apes, although the site of its origin varies, sometimes occurring on
the spine of the scapula. Our Munich anatomist differs from Huxley in
regarding this muscle as “a brilliant proof of the relation of all apes
with each other.” I give this assertion without further comment.
Public-domain text, read in full here on John Shaqi.
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