The muscles which extend between the head, sternum, and clavicle,
together with the muscles of the acromion process of the scapula, make
an external covering to the throat-pouch, which I shall describe
presently. The pectoralis major of the gorilla, as well as that of man,
divides into two portions, one attached to the clavicle, the other
to the cartilages of the true ribs. The former is divided from the
deltoid by a wide interval, filled with connective tissue and fat. But
both portions of the pectoralis major are divided by a tolerably wide
space, into which, in Bischoff’s opinion, the throat-pouch is inserted.
This, however, I do not believe, since that organ would be compressed
and strangulated between the two portions of muscle whenever they were
exercised. It may, however, be supposed that room for an enlargement
of the throat-pouch when the animal is bellowing is afforded by the
existence of these spaces. Bischoff is right in the assertion that
the clavicular portion of the pectoralis major is wanting in the
orang-utan. The upper part of this muscle springs directly from the
sternum. The lower sternal ribs give origin to the pectoralis minor.
The chimpanzee and gibbon display clearly in this muscle the separation
we have mentioned into a clavicular and a sternal portion.
The structure of the pectoralis minor in these apes is full of
interest. In the gorilla it divides into an upper portion of firmer
tissue, less easily separable into digitations, which arises from the
third to the fifth ribs, and a lower portion, separable into three
digitations, of which the upper segment laps considerably over the
lower segment of the upper portion. In the chimpanzee an upper portion
of less firm texture extends from the second to the fourth, and a
lower with three digitations from the fourth to the seventh ribs. This
second lower portion is sometimes absent. I have seen the upper portion
attached to the coracoid process of the scapula, and the lower portion
to the ridge of the greater tuberosity of the humerus. In the orang
an upper portion, separable into three digitations, extends from the
second to the fifth ribs, and is attached to the coracoid process. A
lower portion, also separable into three digitations, extends from the
fifth to the seventh ribs, and is also attached either to the greater
tuberosity of the humerus or to its edge; this latter portion projects
below over the pectoralis major. In the gibbon (_Hylobates albimanus_),
the upper portion starts from the second, the lower from the third to
the fifth ribs. It may here be remarked that the pectoralis minor is in
man also sometimes separable into digitations, which may be connected
both with the coracoid process and with the capsular ligament of the
shoulder-joint. In anthropoids the tendon of insertion of this muscle
is remarkably slender.
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