Anatomy, Artistic; Animals in art; Zoological illustration
The individual whom we have just supposed placed in the attitude of a
quadruped would be able to maintain this position by pressing on the
ground more or less extensive portions of his hands; the whole palm of
the hand may be applied to the ground (Fig. 22); or the fingers
only--that is to say, the phalanges (Fig. 23); or the extremities of
the fingers only--that is to say, the third phalanges (Fig. 24). This
last position, which is certainly difficult to maintain, should here be
regarded rather as theoretical.
We shall meet with each of these modes of support in certain groups of
animals. Thus, the bear, badger, and the majority of rodents, have the
paws applied to the ground by the whole extent of the palmar surface of
the hand, from the wrist to the tips of the fingers. They are therefore
called plantigrade, from the analogy, in this case, of the palm of the
hand to the plantar surface, or sole of the foot.
[Illustration: FIG. 22.--THE HUMAN HAND RESTING FOR ITS WHOLE EXTENT ON
ITS PALMAR SURFACE: LEFT SIDE, EXTERNAL SURFACE.]
In others, such as the lion, tiger, panther, cat, wolf, and fox, the
support is made no longer on the whole extent of the palmar surface, but
on the corresponding surface of the fingers only--the metacarpus is
turned back, and, consequently, the wrist--that is to say, the
carpus--is removed from the ground. These are the digitigrades.
Lastly, the ruminants (sheep, oxen, deer, etc.), and also the pig, ass,
and horse, rest on the third phalanx only. In them not only is the
metacarpus turned back, but also the two first phalanges. The wrist is
very far removed from the ground. In these animals, the third phalanx is
enclosed in a case of horn, a nail (the hoof), and because the support
of the limb is on that nail, the name of unguligrades has been given
them. Nevertheless, as the point of support is on the third phalanx,
which is also known by the name of phalangette, we are of opinion that,
in order to specify definitely, although they walk on their fingers, as
do the digitigrades, the support is provided not by the whole extent of
those appendages, they might receive the name of phalangettigrades.
[Illustration: FIG. 23.--THE HUMAN HAND RESTING ON ITS PHALANGES: LEFT
SIDE, EXTERNAL SURFACE.]
[Illustration: FIG. 24.--THE HUMAN HAND RESTING ON THE TIPS OF SOME OF
ITS THIRD PHALANGES: LEFT SIDE, EXTERNAL VIEW.]
It is necessary among the ruminants to make an exception of the camel
and the llama, which are digitigrades.
Just in proportion as the hand is raised from the ground, as we have
just seen in passing from the plantigrades to the digitigrades and
unguligrades, the number of bones of that region diminishes, the bones
of the forearm coalesce, and the ulna tends to disappear; the hand
becomes less and less suitable for grasping, climbing, or digging, so as
to form an organ exclusively adapted for walking and supporting the
body.
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