Anatomy, Artistic; Animals in art; Zoological illustration
The foot, in animals, as well as in man, is formed of three portions,
which, as we pass from the part which articulates with the leg towards
the terminal extremity, are: the tarsus, the metatarsus, and the toes.
These three portions are the homologues of the carpus, the metacarpus,
and the fingers, which, as we have already seen in the case of the hand,
are the osseous groups which form its skeleton. The tarsus is formed of
short bones, as the carpus is; these are, in man, seven in number. The
bones are arranged in two rows: one, the posterior, formed of two bones
superimposed--the astragalus, by which the tarsus articulates with the
leg, and the calcaneum, which forms the prominence of the heel; and an
anterior row formed of five juxtaposed ones--the cuboid, situated
externally, and the scaphoid internally, in front of which are found the
three cuneiforms. To the tarsus succeeds the metatarsus, whose form
reminds us very much of that of the metacarpals.
With regard to the toes, which we enumerate in proceeding from the most
internal to the most external, they are formed of phalanges, which are
three in number for the four outer toes; but the number is reduced to
two in the case of the first--that is, the so-called great-toe.
The bones of the tarsus are not seven in all animals; they are fewer in
ruminants and solipeds. We already know that, in the latter, the
metacarpals and the digits are equally reduced in number; the same is
the case for the metatarsals and the toes. We will analyze these
differences when dealing with the species individually.
When we studied the anterior limbs, we saw in passing from the
plantigrades to the digitigrades, and finally the ungulates, or
unguligrades, as the hand became hyperextended, the carpus was raised
and more and more removed from the ground. We shall establish the
existence of the same condition in the posterior limbs; in the
plantigrades the tarsus rests on the ground; in the digitigrades it is
removed from it; while in the unguligrades the distance which separates
it from the point of support is still more considerable; and it is,
indeed, necessary to imagine that if these latter were plantigrades,
would occupy the position on the ground which is indicated by Fig. 47.
In veterinary anatomy the tarsus is called the _ham_; a name we adopt
in conformity with usage, but which we cannot but regret, as in human
anatomy the ham is the region of the posterior surface of the knee.
The general arrangement of the region of the digits of the posterior
limbs in birds, presents some points of interest.
We shall merely say with regard to the metatarsus, that it is formed by
a single bone, which in the cock is furnished towards its inferior third
with a pointed process, the _spur_. At the inferior part, there is,
however, found another, which is but very slightly developed, and with
which the first phalanx of the innermost toe articulates.
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