Aspects of plant life; with special reference to the British floraPraeger, R. Lloyd (Robert Lloyd)
Science
Aspects of plant life; with special reference to the British flora
Praeger, R. Lloyd (Robert Lloyd)
Plants -- Great Britain
In contrast to the extreme development in length found in the stems of
climbing plants the extreme reduction of stem found in many plants of
dry places may be referred to. The Crocus, for instance, has an
abbreviated upright stem of which each year’s growth is distended for
the storage of food: one year’s growth dies away as the next enlarges,
so that the well-known bulb-like _corm_ is produced. Compare the
“roots”--really the stem--of Montbretia, in which the annual growths
remain, the result being a knobby structure like a string of onions. In
bulbs reduction in length is carried still farther, the stem forming a
broad cone from the surface of which spring a number of modified leaves,
forming fleshy scales swollen with food material; these surround and
protect the bud, which when it grows produces green leaves and a
terminal flower-shoot; growth is continued by axillary scale-leaved
shoots situated among the scale leaves, which in due course themselves
produce green leaves and flowers. These compact food-charged stems take
up their position well below the ground, out of reach of intense heat or
drought, and during the favourable season send up rapidly into the air
their leaves and flowers, after which they remain dormant till the
following year.
It has been seen that unless a plant is a parasite or saprophyte, using
as food ready-made organic material, it is necessary that it should
possess a sufficient expanse of green (_i.e._, chlorophyll-bearing)
tissue for the purpose of assimilation. This is the essential function
of the leaves; but before leaving the study of stems it should be
pointed out that they usually assist, and sometimes entirely replace,
the leaves as organs of food-manufacture. We have seen how in dry
places--whether physically dry, from direct scarcity of water, or
physiologically dry, owing to reduced activity on the part of the plant
due to unfavourable conditions, such as obtain in cold regions, or on
poisoned ground like salt-marshes or bogs--leaf surface tends to be
reduced, to avoid excessive loss of water. In such plants as the Cacti,
and the Euphorbias which so closely mimic the cactus form, this
reduction is carried to its limit. Leaves are absent, and the stems,
greatly swollen so as to store water, take up the process of
assimilation, and perform it satisfactorily. In more rapid-growing
plants, a sufficient area for assimilation may be obtained by abundant
branching, as in the Gorse, in which leaves are present only in the
seedling stage. In the Brooms (_Genista_) the leaf-development is often
weak, but the stems sometimes make up for this by bearing green
flattened wings. In the Spanish Broom (_G. sagittalis_), a straggling
shrub inhabiting dry places in south-west Europe, the few ovate hairy
leaves, produced in spring, soon fall; but the slender branches bear
several (two to four) broad green wings, which act as
[Illustration: FIG. 20.--GENISTA SAGITTALIS. 1/2.]
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