Aspects of plant life; with special reference to the British floraPraeger, R. Lloyd (Robert Lloyd)
Science
Aspects of plant life; with special reference to the British flora
Praeger, R. Lloyd (Robert Lloyd)
Plants -- Great Britain
leaves, and persist for a couple of years, when they pass away, leaving
slender, round, brown stems. In our native Broom (_Sarothamnus
scoparius_) a similar modification may be observed, though of less
degree. Sometimes stem-structures assume a very leaf-like form, as in
the Butcher’s Broom (_Ruscus aculeatus_), where the ultimate branches
are ovate and quite flat, and might be taken for true leaves but for the
fact that they bear on their surface flowers, and subsequently berries.
The leaves themselves are in this plant reduced to minute scales, and
from their axils these flattened branches spring. In fact, where leaf
reduction takes place, the process of assimilation is often shared in
varying degree by the leaves, the stipules, and the stems. Among our
native plants, as, for instance, in the Leguminosæ and Rosaceæ, the
reader may find for himself many interesting examples for examination.
But the large majority of the Seed Plants bear well-developed leaves, to
which the process of assimilation is practically confined.
LEAVES vary surprisingly in size, shape, and arrangement, features which
are closely related to the characters of the stems which bear them, the
object being the most advantageous display of the chlorophyll in
relation to the light-supply. In general they naturally take the form of
a broad thin blade, protected as may be necessary against extremes of
weather, and guarded against the obvious danger of being dried up by a
thin waterproof covering or cuticle outside the epidermal layer of
cells. In leaves we find the same beauty of mechanical construction as
is seen in stems. The problem is again that of securing maximum
efficiency with minimum expenditure of material. To give as great a
surface as possible, the leaves are as broad and thin as is consistent
with safety, the question of damage by wind being an important
controlling factor. The veins, or vascular bundles, act efficiently as
strengtheners of the thin surface; to prevent tearing at the leaf-edges
the veins are often looped along the margin; while in indented leaves
the extremities of the indentations are strengthened with special
tissue. When one surface of the leaf faces the sky, as in most cases it
does, this surface is strengthened against the weather, and the stomata
are arranged mostly on the lower surface. Where occasionally the leaves
hang normally in a vertical position, as do the mature leaves of the Gum
Trees (_Eucalyptus_), both sides are protected, and the stomata are
borne on the two faces equally. In the Water Lily, again, whose leaves
float, the upper face, which alone is exposed to the air, bears the
stomata, which are present in unusual numbers--nearly 300,000 to the
square inch; the leaf surface is toughened to resist rain and wind, and
waxy to prevent water from lying on it and so interfering with
transpiration. The presence or absence of a leaf-stalk, again, is often
clearly related to the light question. In the Water Lilies the continued
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