Aspects of plant life; with special reference to the British floraPraeger, R. Lloyd (Robert Lloyd)
Science
Aspects of plant life; with special reference to the British flora
Praeger, R. Lloyd (Robert Lloyd)
Plants -- Great Britain
lengthening of the elongated petiole causes the older leaves to float
clear outside of the younger ones. In many biennial herbs, where food is
stored up during the first season in preparation for the flowering
effort in the second, a similar arrangement prevails--note the
leaf-rosettes displayed by Spear Thistle (_Carduus lanceolatus_) and
Herb Robert (_Geranium Robertianum_), as also especially in winter by
perennials like the Dandelion (_Taraxacum officinale_) and Ribwort
(_Plantago lanceolata_). Where stems spread horizontally, as the lower
branches of trees, the leaves are arranged more or less in one
[Illustration: FIG. 21.--AZARA MICROPHYLLA. 1/2.]
plane, in such a manner that overlapping is reduced to a minimum (Fig.
21). This is well seen in horizontal branches of the Elm and other
familiar trees. In the plant chosen for illustration (_Azara
microphylla_, a Chilian shrub), an interesting arrangement obtains. One
of the pair of stipules which subtends each leaf is itself leaf-like,
and stands at an angle, so that a mosaic is formed of true leaves (the
larger ones) and stipules (the smaller alternating ones). On all stems
the leaves are arranged not at haphazard, but according to definite
rules. Sometimes they
[Illustration: FIG. 22.--LEAF OF WEINMANNIA TRICHOSPERMA. 1/1.]
are grouped in circles (_whorls_) at certain points of the stem, as in
the Bedstraws; often in opposite pairs, arranged criss-cross, as in the
Sycamore; most frequently in a series of spirals. The result in all
cases is the same--it allows of as great an interval as possible between
any leaf and the one immediately below or above it, and gives to all an
equal share of light. The indenting of leaves, as in the Sycamore, or
their division into separate segments, as in the Ash and Horse Chestnut,
is of undoubted advantage as allowing light to pass through to lower
layers of leaves; it also materially diminishes the danger arising from
excessive wind-pressure. In the former case there is often a wide space
between the divisions of the leaf; but where this is not required, the
parts of the leaf fit closely together, to secure a maximum of surface.
A particularly pretty example is seen in the Chilian shrub _Weinmannia
trichosperma_ (Fig. 22). Here, to avoid the loss of the area between the
leaflets, the mid rib steps in, developing triangular wings which fill
the spaces. It might be objected that the plant might have saved itself
much trouble by producing, while it was about it, a simple undivided
leaf covering the whole area. It is difficult to answer such
suggestions. Probably the present form of the leaf best meets the
conditions of wind, rain, and light under which it lives. Possibly its
present form is bound up with its ancestral history. “It must be
acknowledged,” says D. H. Scott, “that nothing is more difficult than to
find out why one plant equips itself for the struggle with one device
and another attains the same end in quite a different way.”
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