At the Deathbed of Darwinism: A Series of PapersDennert, Eberhard
Philosophy
At the Deathbed of Darwinism: A Series of Papers
Dennert, Eberhard
Evolution; Natural selection
Haberlandt's experiment, however, had also an indirect consequence that
is of far-reaching importance. He observed that within a few days new
water-secreting organs of an entirely different structure and of
different origin were formed on the leaves that had been sprinkled with
sublimate. Over the bundles of vascular fibres, little knots as large
as a pin head arose in larger numbers out of a tissue underlying the
top layer; out of these the water now oozed every morning. Closer
investigation disclosed the fact that these organs develop only on
young immature leaves where groups of peculiar, perishable gland-hairs
are found; beneath these dead mucous glands the substitute secretive
organs originate in the inner tissue. It is of no importance to state
in what particular cells they originate.
Suffice it to say that they are colorless capillary tubes originating
in various cells; projecting like the hairs of a brush, containing
living protoplasm and evanescent chlorophyll. It is also important
to note that this new organ is immediately connected with the
water-conducting system consisting of bundles of vascular fibres.
Haberlandt furthermore indicates especially that these organs when
viewed in connection with the process of secretion give evidence of an
active vital principle as well as of simple mechanical filtration.
These substitute organs are all indeed well adapted to their purpose
and adequately replace the old secretive organs, but they so easily dry
out and are so little protected that after a week they become parched
and die because wound-cork forms under them. The leaf no longer
produces new hydathodes, but on its lower side it produces growths that
function as vesicles, by means of which it continues to sustain itself.
Haberlandt furthermore records a phenomenon perhaps analogous to this
on the grape-vine, but with this exception the case described by him is
unique. In order to pass any further judgment regarding it, we should
have to ascertain whether the whole phenomenon is not a case of
so-called adaptation; if so, processes should be found in nature,
analogous to the poisoning of the hydathodes in this experiment, which
result in the destruction of the hydathodes so that in consequence the
plant would have gained the power of making good the loss, by means of
the substitute organs. Such processes, however, (even through poisoning
or through parasites) would be very highly improbable. Equally
incredible is the alternative possibility that the new organs would be
produced by the plant not as a substitute but as a supplementary
apparatus when the old ones would not suffice for secretion in case of
very large absorption of water. This also must doubtlessly be rejected,
as Haberlandt has observed.
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