Var. _obtusilobata_, Torrey, Fl. New York, ii., p. 499, t. clx (_Onoclea
obtusilobata_, Schkuhr), is not a permanent variation of the species, but
is based on a not infrequent condition of the plant, in which the pinnæ
of some of the foliaceous fronds become deeply pinnatifid into obovate
segments, which have mostly free veins and imperfectly developed sori.
The indusia appear as little whitish scales on the back of the veins. It
occurs in almost all places where the plant is common, is often produced
from root-stocks which bear also normal fronds, and presents all
gradations from the usual sterile frond to the proper fertile one.
_Ragiopteris onocleoides_ of Presl is founded on a young fertile frond of
this species placed with a sterile one of what Milde judges to be a
monstrous form of _Aspidium Filix-mas_. Maximowicz describes a var.
_interrupta_, from the Amoor region, in which the fertile frond nearly
equals the sterile, and has elongated pinnæ, with remote segments. This
condition is also sometimes seen in American specimens, and is hardly a
true variety.
In an article on “The late Extinct Floras of North America,” which
appeared in Vol. ix of the Annals of the New York Lyceum of Natural
History, in April, 1868, Professor Newberry describes certain fossil
specimens of ferns occurring in Miocene argillaceous limestone at Fort
Union, Dacotah, and refers them with little hesitation to this species. I
have not seen the specimens, but, as similar venation and not very
dissimilar fronds are seen in Woodwardia and Pteris, one may perhaps
doubt the absolute certainty of the identification.
Footnotes
[1]Milde indicates several other unimportant variations; and Hooker &
Baker have as varieties of this species the East Indian _Aspidium
cochleatum_, and _Aspidium elongatum_, from Madeira and the Canary
Islands. The latter they give as occurring also in the southern
United States, evidently supposing it to be the long-lost _A.
Ludovicianum_ of Kunze. For abundant synonymy of _Aspidium Filix-mas_
the student is referred especially to the works of Hooker, Milde,
Mettenius and Moore, as quoted above.
[2]See the “Flora of New York” for some figures of laciniated and forking
fronds.
[3]Prof. Amos Eaton, grandfather of the present writer. Eaton’s “Manual
of Botany” went through eight editions from 1817 to 1841.
[4]I find one or two instances of a slight enlargement of the apex, as if
there were an attempt to form a proliferous bud.
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