The stalks are usually but a few (four to six) inches long, and in the
dried specimens of a brownish straw-color, becoming nearly black at the
base. They bear a few large ferruginous chaffy scales, and are deeply
channelled and furrowed. The fibro-vascular system of the stalk is
altered by contraction in drying, but apparently agrees with Dr. Milde’s
description of _Athyrium_: “There are two oblong peripheric bundles in
the _base_ of the stalk, which, at the base of the lamina, are united
into one of a horse-shoe shape by an arc parallel to the back of the
stalk.” In the middle of a stalk from one of the California specimens I
find two systems of ducts, one on each side of the stalk, and the two
united by a curved and contorted border of firm blackish tissue
(sclerenchyma).
The fronds are from one to two feet long, and from three to six inches
wide. In general shape they are oblong-lanceolate, acuminate, and
slightly narrowed at the base. The texture is softly membranaceous, and
both surfaces are smooth. The primary pinnæ are numerous, the lower ones
gradually farther apart: their shape is lanceolate from a broad base.
They are usually twice pinnatifid, the pinnules being connected by a very
narrow foliaceous border along the midribs. The ultimate segments are
sharply toothed. The fruit-dots are very abundant, and usually are found
on all the pinnæ. They are placed on the back of the free veinlets, and
are apparently devoid of indusium; though Dr. Mettenius has discovered on
young fronds an exceedingly delicate and fugitive indusium, resembling in
some degree that of _Asplenium § Athyrium_. Accordingly, in his later
work, he referred the species to the genus _Asplenium_, placing it next
to _A. Filix-fœmina_. Milde, in his work on the ferns of Europe and
Atlantis, sought to re-establish Athyrium as a genus, and placed this
fern in it, saying “sori ... rotundi, primum breviter oblongi indusio
fugaci minutissimo ciliato instructi.” The spores are ovoid, and
apparently covered with anastomosing raised lines. Those I have examined
are fuscous-brown, but Milde says “sub-nigræ verrucosæ.”
There is a European var. _flexilis_, with very narrow, nearly sessile
fronds, and the pinnæ often deflexed, which has not been observed in
America.
Undoubtedly the greatest resemblance of this fern is to the lady-fern,
_Asplenium Filix-fœmina_; but that species has a very well-developed
indusium, while the minute objects delineated by Mettenius scarcely
deserve the name.
The stalks are clearly continuous with the root-stock; and for this
reason the plant is plainly not a _Polypodium_, whatever else it may
finally be determined to be.
ASPIDIUM FRAGRANS, Swartz.
Fragrant Wood-Fern.
Public-domain text, read in full here on John Shaqi.
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