Lichens were formerly supposed to be a distinct or separate division
of plants. They are now known to be organisms, each species of which
is a constant association of a fungus and an alga. The thallus is
ordinarily made up of fungous mycelium or tissue within which the
imprisoned alga is definitely distributed. The result is a growth
unlike either component. This association of alga and fungus is
usually spoken of as _symbiosis_, or mutually helpful growth, the
alga furnishing some things, the fungus others, and both together
being able to accomplish work that neither could do independently. By
others this union is considered to be a mild form of parasitism, in
which the fungus profits at the expense of the alga. As favourable to
this view, the facts are cited that each component is able to grow
independently, and that under such conditions the algal cells seem to
thrive better than when imprisoned by the fungus.
Lichens propagate by means of _soredia_, which are tiny parts
separated from the body of the thallus, and consisting of one or more
algal cells overgrown with fungus threads. These are readily observed
in many lichens. They also produce spores, usually ascospores, which
are always the product of the fungus element, and which reproduce the
lichen by germinating in the presence of algal cells, to which the
hyphæ immediately cling.
Lichens are found in the most inhospitable places, and, by means of
acids which they secrete, they attack and slowly disintegrate even
the hardest rocks. By making thin sections of the thallus with a
sharp razor and examining under the compound microscope, it is easy
to distinguish the two components in many lichens.
LIVERWORTS
[Illustration: FIG. 288. FIG. 289.
PLANTS OF MARCHANTIA.]
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