The liverworts are peculiar flat green plants usually found on wet
cliffs and in other moist, shady places. They frequently occur
in greenhouses where the soil is kept constantly wet. One of the
commonest liverworts is _Marchantia polymorpha_, two plants of which
are shown in Figs. 288, 289. The plant consists of a ribbon-like
thallus that creeps along the ground, becoming repeatedly forked as
it grows. The end of each branch is always conspicuously notched.
There is a prominent midrib extending along the centre of each branch
of the thallus. On the under side of the thallus, especially along
the midrib, there are numerous rhizoids which serve the purpose of
roots, absorbing nourishment from the earth and holding the plant in
its place. The upper surface of the thallus is divided into minute
rhombic areas that can be seen with the naked eye. Each of these
areas is perforated by a small breathing pore or stomate that leads
into a cavity just beneath the epidermis. This space is surrounded
by chlorophyll-bearing cells, some of which stand in rows from the
bottom of the cavity (Fig. 290). The delicate assimilating tissue is
thus brought in close communication with the outer air through the
pore in the thick, protecting epidermis.
[Illustration: FIG. 290.—SECTION OF THALLUS OF MARCHANTIA. Stomate at
_a_.]
At various points on the midrib are little cups containing small
green bodies. These bodies are buds or _gemmæ_ which are outgrowths
from the cells at the bottom of the cup. They become loosened and are
then dispersed by the rain to other places, where they take root and
grow into new plants.
The most striking organs on the thallus of marchantia are the
peculiar stalked bodies shown in Figs. 288, 289. These are termed
_archegoniophores_ and _antheridiophores_ or _receptacles_. Their
structure and function are very interesting, but their parts are so
minute that they can be studied only with the aid of a microscope
magnifying from 100 to 400 times. Enlarged drawings will guide the
pupil.
[Illustration: FIG. 291.—SECTION THROUGH ANTHERIDIOPHORE OF
MARCHANTIA, showing antheridia. One antheridium more magnified.]
The _antheridiophores_ are fleshy, lobed disks borne on short stalks
(Fig. 291). The upper surface of the disk shows openings scarcely
visible to the naked eye. However, a section of the disk, such as is
drawn in Fig. 291, shows that the pores lead into oblong cavities
in the receptacle. From the base of each cavity there arises a
thick, club-shaped body, the _antheridium_. Within the antheridium
are formed many sperm-cells which are capable of swimming about in
water by means of long lashes or cilia attached to them. When the
antheridium is mature, it bursts and allows the ciliated sperm cells
to escape.
[Illustration: FIG. 292.—ARCHEGONIUM OF MARCHANTIA.]
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