If we have followed carefully the development of marchantia, the
study of one of the mosses will be comparatively easy. The mosses
are more familiar plants than the liverworts. They grow on trees,
stones, and on the soil in both wet and dry places. One of the
common larger mosses, known as _Polytrichum commune_, may serve as
an example, Fig. 295. This plant grows on rather dry knolls, mostly
in the borders of open woods, where it forms large beds. In dry
weather these beds have a reddish brown appearance, but when moist
they form beautiful green cushions. This colour is due, in the first
instance, to the colour of the old stems and leaves, and, in the
second instance, to the peculiar action of the green living leaves
under the influence of changing moisture-conditions. The inner or
upper surface of the leaf is covered with thin, longitudinal ridges
of delicate cells which contain chlorophyll. These cells are shown in
cross-section in Fig. 296, as dots or granules. All the other tissue
of the leaf consists of thick-walled, corky cells which do not allow
moisture to penetrate. When the air is moist the green leaves spread
out, exposing the chlorophyll cells to the air, but in dry weather
the margins of the leaves roll inward, and the leaves fold closely
against the stem, thus protecting the delicate assimilating tissue.
[Illustration: FIG. 296.—SECTION OF LEAF OF POLYTRICHUM COMMUNE.]
The _antheridia_ and _archegonia_ of polytrichum are borne in groups
at the ends of the branches on different plants (many mosses bear
both organs on the same branch). They are surrounded by involucres
of characteristic leaves termed _perichætia_ or _perichætal leaves_.
Multicellular hairs known as _paraphyses_ are scattered among the
archegonia and antheridia. The involucres with the organs borne
within them are called _receptacles_, or, less appropriately, “moss
flowers.” As in marchantia, the organs are very minute and must be
highly magnified to be studied.
[Illustration: FIG. 297.—SECTION THROUGH A RECEPTACLE OF POLYTRICHUM
COMMUNE, showing paraphyses and antheridia. ]
The antheridia are borne in broad cup-like receptacles on the
antheridial plants (Fig. 297). They are much like the antheridia of
marchantia, but they stand free among the paraphyses and are not
sunk in cavities. At maturity they burst and allow the sperm-cells
or _spermatozoids_ to escape. In polytrichum, when the receptacles
have fulfilled their function, the stem continues to grow from the
centre of the cup (_m_, Fig. 295). The archegonia are borne in other
receptacles on different plants. They are like the archegonia of
marchantia except that they stand erect on the end of the branch.
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