The _sporogonium_ which develops from the fertilized egg is shown
in _a_, _b_, Fig. 295. It consists of a long, brown stalk bearing
the spore-case at its summit. The base of the stalk is imbedded
in the end of the moss stem by which it is nourished. The capsule
is entirely inclosed by a hairy cap, the _calyptra_, _b_. The
calyptra is really the remnant of the archegonium, which, for
a time, increases in size to accommodate and protect the young
growing capsule. It is finally torn loose and carried up on the
spore-case. The mouth of the capsule is closed by a circular lid, the
_operculum_, having a conical projection at the centre.
The operculum soon drops, or it may be removed, displaying a fringe
of sixty-four teeth guarding the mouth of the capsule. This ring of
teeth is known as the _peristome_. In most mosses the teeth exhibit
peculiar hygroscopic movements; _i.e._ when moist they bend outwards,
and upon drying curve in toward the mouth of the capsule. This
motion, it will be seen, serves to disperse these spores gradually
over a long period of time.
Not the entire capsule is filled with spores. There are no elaters,
but the centre of the capsule is occupied by a columnar strand of
tissue, the _columella_, which expands at the mouth into a thin,
membranous disk, closing the entire mouth of the capsule except the
narrow annular chink guarded by the teeth. In this moss the points of
the teeth are attached to the margin of the membrane, allowing the
spores to sift out through the spaces between them.
When the spores germinate they form a green, branched thread, the
_protonema_. This gives rise directly to moss plants, which appear
as little buds on the thread. When the moss plants have sent their
little rhizoids into the earth, the protonema dies, for it is no
longer necessary for the support of the little plants, and the moss
plants grow independently.
* * * * *
[Illustration: FIG. 298.—FUNARIA HYGROSCOPICA. ]
_Funaria_ is a moss very common on damp, open soil. It forms green
patches of small fine leaves from which arise long brown stalks
terminated by curved capsules (Fig. 298). The structure is similar to
that of polytrichum, except the absence of plates on the under side
of the leaves, the continuous growth of the stem, the curved capsule,
double peristome, monœcious rather than diœcious receptacles, and
nearly glabrous unsymmetrical calyptra.
EQUISETUMS, OR HORSETAILS (Pteridophyta)
There are about twenty-five species of equisetum, constituting the
only genus of the unique family _Equisetaceæ_. Among these _E.
arvense_ (Fig. 299) is common on clayey and sandy soils.
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