In this species the work of nutrition and that of spore production
are performed by separate shoots from an underground rhizome. The
fertile branches appear early in spring. The stem, which is 3
to 6 inches high, consists of a number of cylindrical, furrowed
internodes, each sheathed at the base by a circle of scale leaves.
The shoots are of a pale yellow colour. They contain no chlorophyll,
and are nourished by the food stored in the rhizome (Fig. 299).
The spores are formed on specially developed fertile leaves or
_sporophylls_ which are collected into a spike or cone at the end of
the stalk (_a_, Fig. 299). A single sporophyll is shown at _b_. It
consists of a short stalk expanded into a broad, mushroom-like head.
Several large _sporangia_ are borne on its under side. The spores
formed in the sporangia are very interesting and beautiful objects
when examined under the microscope (× about 200). They are spherical,
green bodies, each surrounded by two spiral bands attached to the
spore at their intersection, _s_. These bands exhibit hygroscopic
movements by means of which the spores become entangled, and are held
together. This is of advantage to the plant, as we shall see. All
the spores are alike, but some of the _prothallia_ grow to a greater
size than the others. The large prothallia produce only _archegonia_
while the smaller ones produce _antheridia_. Both these organs are
much like those of the ferns, and fertilization is accomplished in
the same way. Since the prothallia are usually diœcious, the special
advantage of the spiral bands, holding the spores together so that
both kinds of prothallia may be in close proximity, will be easily
understood. As in the fern, the fertilized egg-cell develops into an
equisetum plant.
[Illustration: FIG. 299.—EQUISETUM ARVENSE.
_st_, sterile shoot;
_f_, fertile shoot showing the spike at _a_;
_b_, sporophyll, with sporangia;
_s_, spore.]
The sterile shoots (_st_, Fig. 299) appear much later in the season.
They give rise to repeated whorls of angular or furrowed branches.
The leaves are very much reduced scales, situated at the internodes.
The stems are provided with chlorophyll and act as assimilating
tissue, nourishing the rhizome and the fertile shoots. Nutriment is
also stored in special tubers developed on the rhizome.
Other species of equisetum have only one kind of shoot—a tall, hard,
leafless, green shoot with the spike at its summit. Equisetum stems
are full of silex, and they are sometimes used for scouring floors
and utensils; hence the common name “scouring rush.”
ISOËTES (Pteridophyta)
_Isoëtes_ or quillwort is usually found in water or damp soil on the
edges of ponds and lakes. The general habit of the plant is seen
in Fig. 300, _a_. It consists of a short, perennial stem bearing
numerous erect, quill-like leaves with broad sheathing bases. The
plants are commonly mistaken for young grasses.
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