_Chlorophyll absorbs certain light rays, and the energy thus directly
or indirectly obtained is used by the living matter in uniting the
carbon dioxide absorbed from the air with some of the water brought
up from the roots. The ultimate result usually is starch._ The
process is obscure, but sugar is generally one step; and our first
definite knowledge of the product begins when starch is deposited
in the leaves. The process of using the carbon dioxide of the air
has been known as carbon assimilation, but the term now most used is
=photosynthesis= (from two Greek words meaning _light_ and _placing
together_.)
=Starch and Sugar.=—_All starch is composed of carbon, hydrogen,
and oxygen_ (C_{6}H_{10}O_{5})_{_n_}. The sugars and the substance
of cell walls are very similar to it in composition. All these
substances are called =carbohydrates=. In making fruit sugar from the
carbon and oxygen of carbon dioxide and from the hydrogen and oxygen
of the water, _there is a surplus of oxygen_ (6 parts CO_{2} + 6
parts H_{2}O = C_{6}H_{12}O_{6} + 6 O_{2}). It is this oxygen that is
given off into the air during sunlight.
=Digestion.=—_Starch is in the form of insoluble granules. When such
food material is carried from one part of the plant to another for
purposes of growth or storage, it is made soluble before it can be
transported._ When this starchy material is transferred from place to
place, it is usually changed into sugar by the action of a diastase.
_This is a process of_ =digestion=. It is much like the change of
starchy foodstuffs to sugary foods effected by the saliva.
[Illustration: FIG. 118.—TRUNK GIRDLED BY A WIRE. See Fig. 85.]
=Distribution of the Digested Food.=—After being changed to the
soluble form, _this material is ready to be used in growth_, either
in the leaf, in the stem, or in the roots. With other more complex
products it is then _distributed throughout all the growing parts of
the plant_; and when passing down to the root, it seems to pass more
readily through the _inner bark_, in plants which have a definite
bark. This gradual downward diffusion through the inner bark of
materials suitable for growth is the process referred to when the
“descent of sap” is mentioned. Starch and other products are often
_stored in one growing season to be used in the next season_. If a
tree is constricted or strangled by a wire around its trunk (Fig.
118), the digested food cannot readily pass down and it is stored
above the girdle, causing an enlargement.
Public-domain text, read in full here on John Shaqi.
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