=Transpiration.=—Much more water is absorbed by the roots than is
used in growth, _and this surplus water passes from the leaves into
the atmosphere by an evaporation process known as_ =transpiration=.
Transpiration takes place more abundantly from the under surfaces
of leaves, and through the pores or stomates. A sunflower plant of
the height of a man, during an active period of growth, gives off a
quart of water per day. A large oak tree may transpire 150 gallons
per day during the summer. For every ounce of dry matter produced, it
is estimated that 15 to 25 pounds of water usually passes through the
plant.
_When the roots fail to supply to the plant sufficient water
to equalize that transpired by the leaves_, =the plant wilts=.
Transpiration from the leaves and delicate shoots is increased
by all the conditions which increase evaporation, such as higher
temperature, dry air, or wind. The stomata open and close, tending to
regulate transpiration as the varying conditions of the atmosphere
affect the moisture content of the plant. However, in periods of
drought or of very hot weather, and especially during a hot wind, the
closing of these stomates cannot sufficiently prevent evaporation.
The roots may be very active and yet fail to absorb sufficient
moisture to equalize that given off by the leaves. The plant shows
the effect (how?). On a hot dry day, note how the leaves of corn
“roll” towards afternoon. Note how fresh and vigorous the same leaves
appear early the following morning. Any injury to the roots, such as
a bruise, or exposure to heat, drought, or cold may cause the plant
to wilt.
Water is forced up by =root pressure= or =sap pressure=. (Exercise
99.) Some of the dew on the grass in the morning may be the water
forced up by the roots; some of it is the condensed vapour of the air.
_The wilting of a plant is due to the loss of water from the cells._
The cell walls are soft, and collapse. A toy balloon will not stand
alone until it is inflated with air or liquid. In the woody parts of
the plant the cell walls may be stiff enough to support themselves,
even though the cell is empty. Measure the contraction due to wilting
and drying by tracing a fresh leaf on page of notebook, and then
tracing the same leaf after it has been dried between papers. The
softer the leaf, the greater will be the contraction.
=Storage.=—We have said that starch may be stored in twigs to be used
the following year. The very early flowers on fruit trees, especially
those that come before the leaves, and those that come from bulbs,
as crocuses and tulips, are supported by the starch or other food
that was organized the year before. Some plants have very special
storage reservoirs, as the potato, in this case being a thickened
stem although growing underground. (Why a thickened stem? p. 84.) It
is well to make the starch test on winter twigs and on all kinds of
thickened parts, as tubers and bulbs.
Public-domain text, read in full here on John Shaqi.
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