The simplest kind of fruit is a _ripened 1-loculed ovary_. The first
stage in complexity is a ripened _2- or many-loculed ovary_. Very
complex forms may arise by the _attachment of other parts to the
ovary_. Sometimes the style persists and becomes a beak (mustard
pods, dentaria, Fig. 224), or a tail as in clematis; or the calyx
may be attached to the ovary; or the ovary may be embedded in the
receptacle, and ovary and receptacle together constitute the fruit:
or an involucre may become a part of the fruit, as possibly in the
walnut and the hickory (Fig. 225), and the cup of the acorn (Fig.
226). The chestnut and the beech bear a prickly involucre, but the
nuts, or true fruits, are not grown fast to it, and the involucre
can scarcely be called a part of the fruit. A ripened ovary is a
=pericarp=. A pericarp to which other parts adhere has been called an
accessory or reënforced fruit. (Page 169.)
[Illustration: FIG. 225.—HICKORY-NUT. The nut is the fruit, contained
in a husk. ]
[Illustration: FIG. 226.—LIVE-OAK ACORN. The fruit is the “seed”
part; the involucre is the “cup.” ]
Some fruits are =dehiscent=, or split open at maturity and liberate
the seeds; others are =indehiscent=, or do not open. A dehiscent
pericarp is called a =pod=. The parts into which such a pod breaks
or splits are known as =valves=. In indehiscent fruits the seed is
liberated by the decay of the envelope, or by the rupturing of the
envelope by the germinating seed. Indehiscent winged pericarps are
known as =samaras= or =key fruits=. Maple (Fig. 227), elm (Fig. 228),
and ash (Fig. 93) are examples.
[Illustration: FIG. 227.—KEY OF SUGAR MAPLE.]
[Illustration: FIG. 228.—KEY OF COMMON AMERICAN ELM.]
=Pericarps.=—The simplest pericarp is a dry, one-seeded, indehiscent
body. It is known as an =akene=. A head of akenes is shown in Fig.
229, and the structure is explained in Fig. 230. Akenes may be seen
in buttercup, hepatica, anemone, smartweed, buckwheat.
[Illustration: FIG. 229.—AKENES OF BUTTERCUP.]
[Illustration: FIG. 230.—AKENES OF BUTTERCUP, one in longitudinal
section. ]
A 1-loculed pericarp which dehisces along the front edge (that is,
the inner edge, next the centre of the flower) is a =follicle=. The
fruit of the larkspur (Fig. 231) is a follicle. There are usually
five of these fruits (sometimes three or four) in each larkspur
flower, each pistil ripening into a follicle. If these pistils were
united, a single compound pistil would be formed. Columbine, peony,
ninebark, milkweed, also have follicles.
[Illustration: FIG. 231.—FOLLICLE OF LARKSPUR.]
[Illustration: FIG. 232.—A BEAN POD.]
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