=Movements: the Water-tube System.=—The water, which is filtered through
the perforated madreporite, is needed to supply a _system of canals_
(Fig. 56). The madreporite opens into a canal called the _stone canal_,
the wall of which is hardened by the same kind of material as that found
in the skin. The stone canal leads to the _ring canal_ which surrounds
the mouth (Fig. 56). The ring canal sends _radial canals_ into each ray
to supply the double row of _tube feet_ found in the groove at the lower
side of each ray (Fig. 57). Because of their arrangement in rows, the
feet are also called _ambulacral_ feet (Latin _ambulacra_, “forest
walks”). There is a water holder (_ampulla_), or muscular water bulb at
the base of each tube foot (Fig. 58). These contract and force the water
into the tube feet and extend them. The cuplike ends of the tubes cling
to the ground by suction. The feet contain delicate muscles by which
they contract and shorten. Thus the animal pulls itself slowly along,
hundreds of feet acting together. The tube feet, for their own
protection, may contract and retire into the groove, the water which
extended them being sent back into the ampulla. This system of water
vessels (or water-vascular system) of the echinodermata is
characteristic of them; _i.e._ is not found elsewhere in the animal
kingdom. The grooves and the plates on each side of them occupy the
_ambulacral areas_. The rows of spines on each side of the grooves are
freely movable. (What advantage?) The spines on the aboral surface are
not freely movable.
[Illustration:
FIG. 57.—Starfish, from below; tube feet extended.
]
[Illustration:
FIG. 58.—SECTION OF ONE RAY and central portion of starfish.
_f_{1}_, _f_{2}_, _f_{3}_, tube feet more or less extended; _au_, eye
spot; _k_, gills; _da_, stomach; _m_, madreporite; _st_, stone
canal; _p_, ampulla; _ei_, ovary.
]
=Respiration.=—The _system of water vessels serves the additional
purpose_ of bringing water containing oxygen into contact with various
parts of the body, and the starfish was formerly thought to have no
special respiratory organs. However, there are holes in the aboral wall
through which the folds of the delicate lining membrane protrude. These
are now supposed to be _gills_ (_k_, Fig. 58).
[Illustration:
FIG. 59.—Starfish eating a sea snail.
_b_, stomach everted.
]
=The nervous system= is so close to the aboral surface that much of it
is visible without dissection. Its chief parts are a _nerve ring_ around
the mouth, which sends off a _branch_ along each ray. These branches may
be seen by separating the rows of tube feet. They end in a pigmented
cell at the end of each ray called the _eye-spot._
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