Birds and Nature Vol. 09 No. 2 [February 1901]Various
History
Birds and Nature Vol. 09 No. 2 [February 1901]
Various
Birds -- Periodicals; Natural history -- Periodicals
The mouth is placed in the lower plane of the head and is recognized
externally as a simple slit. Inside of the mouth is placed one of the
most wonderful dental apparatuses known to science. This is called the
radula, odontophore or tooth-bearer, and is a belt of chitinous,
transparent, yellowish or colorless material, its upper surface being
armed with numerous siliceous teeth arranged in longitudinal and
parallel rows. The radula is placed in an organ called the buccal sac
and occupies a position in the sac analogous to that of the tongue in a
cat or dog, viz., on the floor of the mouth. It is formed from a layer
of cells in the posterior part of the buccal sac, called the radula sac,
and new teeth are constantly forming here to take the place of those
which have become worn by use. The whole radula rests upon a cartilage,
is strongly fastened at the anterior end, and is brought down between
the two fleshy lips of the mouth where it performs a backward and
forward movement, thus rasping off with the sharp teeth particles of
food which have been cut into small pieces by the horny jaw. During this
process the morsel of food is pressed against the top or roof of the
mouth. The jaw is placed in the upper part of the mouth in front of the
radula, and is frequently armed with ribs to aid in cutting or biting
off pieces of food, as leaves or vegetables.
As before remarked, the radula is made up of parallel rows of teeth, the
whole area being usually divided into five longitudinal rows, each row
differing from the one next to it. We have first a central row, on each
side of this a lateral row and finally a marginal row. Each tooth in
each row is made up of different parts, a basal part attached to the
radula belt and an upper part which is turned over or reflexed and bent
backward so as to tear off food particles by a backward movement of the
whole apparatus. This diversity of form in the teeth has led
conchologists to adopt a tooth formula similar to that adopted for
vertebrate animals, so that the teeth of different species can be
compared and the animals classified thereby. Thus each tooth has certain
prominences called cusps, which vary in size, number and position, and
serve admirably to describe the different groups of snails. All the
mollusca, except the bivalves, are provided with this radula.
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