The root-system of _Acanthyllis_ offered some points of interest
and a short study of it was made in the field. All of the specimens
whose roots were examined were growing in the habitat above referred
to and within a meter of one another. The leading results of the
observation are as follows: The largest specimen studied possessed a
tap-root 3 cm. in diameter at the crown. Growing rapidly smaller as
it ran downward, the root gave off four laterals, of which a portion
dipped at an acute angle to a depth of 20 to 30 cm., sending off
branches by the way. One of the largest of the laterals was traced
75 cm. and where left was 2 cm. beneath the surface. The branches,
at least of the main laterals, that is, the roots of the tertiary
order, for the most of their course ran thus near the surface of
the ground. One of the leading laterals was followed to the base of
a neighboring specimen of quedad, where it lay close to the crown
of the main root. The depth of the penetration of the tap-root
of this specimen was not learned. The tap-root of a neighboring
specimen ran directly downward 20 cm.; then, turning sharply,
it extended in a horizontal direction for a distance of 70 cm. As
the large laterals were wholly lacking on this plant, the tap-root
was the entire system—surely an anomalous condition. On a third
plant the tap-root penetrated the ground about 4 cm. only, after
which it turned and ran the rest of its course within 4 cm. of the
surface. The more superficial of the roots of a fourth specimen were
found to extend to the base of the last plant mentioned. Thus it was
found that the root-system of _Acanthyllis_, as growing naturally,
extends both widely and deeply for a considerable distance, and that
it is flexible to a degree; in short, is generalized[8] and closely
resembles that of certain species of the Tucson region, particularly
_Covillea tridentata_, which grows under similar conditions.
The root-systems of three or four other species were also
examined. Of these _Zollikoferia spinosa_ and _Artemisia campestris_
were growing in a little hollow in the plain close by the habitat
of _Acanthyllis_. _Zollikoferia_ has a very close habit of growth
with dichotomous branching (compare figs. 5 and 7). When dry the
branches are slender and of a woody hardness. The root-system of
this species is characterized by a pronounced tap-root and by the
absence of large laterals, at least near the surface. The leading
feature of the root-system is, in short, its tap-root. A root-system
of a similar type was found in _Artemisia herba-alba_.
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