Some stems accomplish their purpose not by holding fast to a support in
the air, but by creeping along the ground, as in the running blackberry,
and often in the Virginia creeper. The purpose is the same, and, as if
to confirm it, a few otherwise quite prostrate vines have their tips
turned upward to the light, notably in the case of the creeping
speedwell.
In certain plants the stem may assume curious forms due to special
conditions under which they live and to which adjustment is necessary
for the plant’s existence. In deserts, for instance, the cactus produces
practically no leaves (Figure 10) and the green stem performs not only
the function of leaves but acts as a storage for water. Where water is
scarce this is of tremendous advantage, a single cactus having been
known to store up 125 gallons. A similar habit of the cactuslike spurges
in South Africa gives as weird an atmosphere to parts of their
landscapes as we find in Arizona. It is as if the stem of such plants,
being unable to push its leaves (it has none) up into the light, takes
over some of the functions of leaves and makes up the deficiency by
adopting other methods
[Illustration: FIG. 10.--COMMON PRICKLY PEAR CACTUS (_Opuntia Opuntia_)
Native along the Atlantic Coast. The green joints of the stem function
as leaves and store water.]
to secure the plant’s survival. Other stems, looking and acting like
leaves, reveal their true nature by producing buds, and the curious
feature of the common butcher’s-broom (Figure 11), often colored scarlet
for Christmas decorations, bearing flowers from the middle of what is
apparently a leaf, but is actually a modified stem, is explained by this
ability of stems to modify their habits to suit conditions. The
butcher’s-broom is an inhabitant of dry regions along the Mediterranean,
where a reduction or
[Illustration: FIG. 11.--BUTCHER’S-BROOM
(_Ruscus aculeatus_)
Note leaflike stems with flowers arising from the center.]
absence of leaf surface is a decided advantage. In many partly desert or
dry regions this production of leaflike stems or branches is common, an
excellent garden example being asparagus, which came originally from
Europe and the feathery growth of which is all stem. In Tasmania a kind
of yew tree produces no leaves, all the foliage being modified stem,
which is true of many kinds of spurge in the West Indies, where an
almost impenetrable scrub is largely made up of a shrub which is
apparently covered with leaves, all actually part of the branches and
stems.
[Illustration: FIG. 12.--DUCKWEED
The smallest known flowering plant, with _no_ leaves and tiny leaflike
stems floating on the surface. Flowers are borne from the margin of the
stem. (Eight times natural size).]
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