Professor Huxley, about thirty years ago, proposed to classify birds
by the form of the bones of the palate. In the skull of the Hawk, it
will be seen that two bones lying in the front region of the palate
are fused with each other in the middle line, and to the type of skull
which is thus characterised the name ‘desmognathous’ was given. It is
found not only in the Hawks, but in a quantity of other birds; for
instance, in the Stork tribe, and in the Hornbills and Toucans. The
second form of skull distinguishes the gallinaceous birds; in them the
two maxillo-palatines remain unconnected, and the palate is therefore
in a way cleft; this is termed the ‘schizognathous’ skull. In the
finch tribe there is a slight modification of this, called, from the
Greek word for a finch, ‘ægithognathous.’ In these birds a median bone,
called the vomer, from the fact that the bone to which it corresponds
in the human skull is shaped somewhat like a plough-share, is truncated
in front, instead of tapering, as it does in the schizognathous skull
of the common fowl. There is a fourth variety, which marks out the
Ostrich tribe and the American Tinamous, in which the two pairs of
bones called the pterygoids and palatines do not, as they do in the
types of skull that have been hitherto considered, reach the middle
line of the skull, but are kept off from it by the vomers, which
extend backwards. The term ‘dromæognathous,’ or emu-like, is applied
to this form of skull. If the back of any bird’s skull be examined,
it will be noticed that just below the great hole or foramen, through
which the medulla passes to join the spinal cord in the canal of the
vertebral column, is a rounded, rather kidney-shaped boss. This is the
occipital condyle, by means of which the skull articulates with the
first vertebra. If you look at the same region in a mammal, you will
find that there are two of these, one on each side, though also below
the foramen magnum. This is one of the many points of structure that
distinguish a bird from a mammal and ally it to the reptiles; but it
must be remembered that in some reptiles there is a commencing division
of the single condyle into two.
The Vertebral Column.
Public-domain text, read in full here on John Shaqi.
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