We must enter into the matter of wing a little more closely--it is so
important a feature of bird organisation. The wing, of course, although
it performs so different a _rôle_, is the exact equivalent of
the fore limb of mammals. We can easily recognise precisely the same
bones, though they are diminished in number, and often of a different
form. It will be noticed that in each case we can distinguish the
three bones forming the arm, and which are known as the humerus, the
radius, and ulna. The rest of the limb in the bird is not quite so
obviously like the hand of the mammal; but a little attention will show
that it is constructed upon a perfectly similar plan. The flexible
wrist of the mammal is made up of many small bones; the hand itself
is made up of a larger series still, of which those nearest to the
wrist are technically termed the metacarpals, and those which follow,
the phalanges. In many mammals there are five fingers; but there are
many which have less, and the extreme is reached in the horse, which
has to put up with a single finger and small rudiments of two others.
Now the bird is better off in the way of fingers than the horse, as it
has three fairly well-developed fingers, or rather two well developed
and one less perfect. The shortest finger corresponds to the thumb of
our hand. It is more freely movable than the others. The metacarpal
bones of the second and third fingers are firmly welded together, and
are long; each finger (as will be seen from a look at fig. 1, p. 4)
has one or two phalanges, as the case may be. Now in mammals the end
phalanx of each finger is tipped with a nail, or with a hoof. The
powerful claws of the tiger, used for tearing, and the solid hoof of
the ox or horse, upon which the creature walks, are one and the same
thing. It might be supposed that the hand of the bird, which is not
an organ of offence or meant to walk with, might be shorn of these
appendages. But this is not the case: every bird has at least two nails
(fig. 9), of a long and rather claw-like form when well developed,
and sometimes three nails, that is, one to each of its fingers. It
looks, therefore, very much as if the wing of the bird had been formed
out of a limb that was once an organ for climbing or walking with.
There is a curious bird, found in British Guiana, which is known as
the Hoatzin (figs. 9, 11). In the very young nestlings of the hoatzin
the claws of the fingers are so conspicuous that they are actually
used by the callow chick to climb with, before the feathers of the
wings have grown sufficiently to enable them to use their wings in the
proper way in which a bird should; it has been said also, that other
birds scramble about and use their claws when they are young. In the
case of the hoatzin, it is stated that the thumb and the first finger
can be brought together so as to lay hold definitely of an object. A
very important thing to notice about the wing bones is that they are
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