too far from the subject at hand; but anyone who would pursue this
further can consult Professor Huxley’s ‘Elementary Physiology,’ or any
other elementary text-book upon physiology. When a mammal--a human
being, for example--breathes certain muscles are called into play. If
a person is watched, it will be seen that the chest expands during
inspiration, and that its calibre diminishes during expiration. What
happens is this. The lungs are contained in a cavity which contains no
air. This cavity can be increased in size in two directions. When the
ribs are moved out--which they can be by the movements of the muscles
called intercostal, which lie between them--the cavity of the chest
from before backwards is evidently enlarged. On the other hand there
is the diaphragm, which we have already spoken of as bounding the
chest cavity below. Now this diaphragm is muscular, with a tendinous
centre. When the muscles contract, like all muscles do, the surface
of the diaphragm, which was before rather convex towards the chest
cavity, becomes more flat; hence the cavity lying above it, _i.e._
the chest cavity, becomes larger in a downward direction also. When
it is increased in this way by the action of the two separate sets of
muscles, some space--more space than before--is left between its walls
and the lungs which lie within it; it follows, therefore, that, as
there is no air in the cavity, the pressure of air outside the body
forces more air into the lungs, because there is no counterbalancing
pressure to prevent this. The principle is the same in the bird,
but the details are different. If you will turn again to the bird’s
skeleton, you will see that the backbone and ribs and sternum form a
bony box, which is jointed in the middle; this acts precisely like a
pair of bellows: the bones at top and bottom represent the wood, and
the soft intervening leather of the bellows is represented by the
muscles which lie between, and which connect the sternum with the
abdomen and with the ribs. When these muscles contract, the sternum
is obviously brought nearer to the backbone, and air is expelled
from the inside; when they are relaxed, a vacuum is created and air
rushes in. The air-spaces, then, are really ramified tags of lung
which have no blood-vessels in their walls, and are therefore not
meant for respiration, but serve as reservoirs of air, lightening
the body of the creature. It is curious that birds are not the only
animals which possess expansions of lung that are apparently useless
for breathing purposes. The lungs of the Chameleon have quite similar
sacs appended to them. There is, it is true, no such complicated a
ramification as that which we find in the bird, but still there is no
doubt that the structure is of the same nature. It looks almost like
a first step in the path towards a bird. Very possibly the extinct
Pterodactyles, which flew through the woods of the middle ages of the
earth, had bodies lightened in the same or a similar way; for we know
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