The elder Darwin writes in a note that “he is acquainted with a philosopher who thinks it not impossible that the first insects were the anthers or stigmas of flowers, which by some means, etc. etc.” This is mere speculation, not a definite theory, and though the passage above as quoted by the “Savoyard” certainly does contain the germ of Darwin’s theory, what is it more than the crudest and most unshapen germ? And in what conceivable way does this discovery of the egg invalidate the excellence of the chicken? Was there ever a great theory yet which was not more or less developed from previous speculations which were all to a certain extent wrong, and all ridiculed, perhaps not undeservedly, at the time of their appearance? There is a wide difference between a speculation and a theory. A speculation involves the notion of a man climbing into a lofty position, and descrying a somewhat remote object which he cannot fully make out. A theory implies that the theorist has looked long and steadfastly till he is clear in his own mind concerning the nature of the thing which he is beholding. I submit that the “Savoyard” has unfairly made use of the failure of certain speculations in order to show that a distinct theory is untenable. Let it be granted that Darwin’s theory has been foreshadowed by numerous previous writers. Grant the “Savoyard” his Giordano Bruno, and give full weight to the barrel-organ in a neighbouring settlement, I would still ask, has the theory of natural development of species ever been placed in anything approaching its present clear and connected form before the appearance of Mr. Darwin’s book? Has it ever received the full attention of the scientific world as a duly organised theory, one presented in a tangible shape and demanding investigation, as the conclusion arrived at by a man of known scientific attainments after years of patient toil? The upshot of the barrel-organs article was to answer this question in the affirmative and to pooh-pooh all further discussion. It would be mere presumption on my part either to attack or defend Darwin, but my indignation was roused at seeing him misrepresented and treated disdainfully. I would wish, too, that the “Savoyard” would have condescended to notice that little matter of the bear. I have searched my copy of Darwin again and again to find anything relating to the subject except what I have quoted in my previous letter. I am, Sir, your obedient servant, A. M. Darwin on Species [From the _Press_, April 11th, 1863.] To the Editor of the _Press_.