Closely pressed against the hinder wall of this bulla is a sort of
bony clamp, which seems to keep the bulla in its place, and running
obliquely along the surface of the swelling is an indistinct groove,
corresponding to which, in the interior of the drum, is a bony wall,
dividing the drum cavity into an inner and an outer compartment, these
two divisions being formed from separate bones, as an examination of a
very young skull will show.
The almost globular form and great relative size of the _bulla
tympani_; the absence of any distinct bony passage leading from its
cavity to the interior, the opening being quite flush with the wall
of the drum; and the division of the cavity into two parts by a bony
partition, are all very important as distinctive characters of the Cat
family, and also, with lesser modifications, of the whole Æluroid group.
[Illustration: UNDER VIEW OF LION’S SKULL.
The letters have the same significance as in the side view.]
The power of retracting the claws, so characteristic a feature of all
the true Cats (which are, without exception, digitigrade), is brought
about by certain peculiarities of structure of the last two joints of
the toes. Of the three _phalanges_, or bones which make up the skeleton
of the toe, the first, or that nearest to the wrist or ankle, is of
the ordinary shape: about three times as long as broad, with a regular
cylindrical shaft, and pulley-like ends, for articulation with the bone
to which it is joined. The second, or middle phalanx, is pretty much
like the first, except that its shaft is scooped out on one side, so
as to make a greater distance between it and the corresponding bone of
the next toe than there would otherwise be. The third and last joint,
called the _ungual phalanx_, from the fact of its supporting the
claw, has the regular pulley-surface to articulate with the preceding
joint, but its farther end is strongly curved downwards and pointed
at the end; it has, in fact, the shape of the horny talon of which it
forms the supporting core. Further support is afforded to the claw by
an outgrowth of the phalanx, which commences near its articular end,
and grows over the end of the claw like a sort of hood, thus giving
the ungual phalanx of the Cat a most peculiar and unmistakable shape.
Between the upper surfaces of the last phalanx and the last but one
passes a strong and very elastic ligament, which so pulls upon the
ungual phalanx as to bend it on its predecessor, and so cause the two
to be almost parallel, the hood of the claw-bearing bone being received
between the preceding joint of its own toe and that of the next; hence
the scooping out of the middle phalanges. Thus, by the action of this
ligament, the claw under ordinary circumstances is pulled back within
its covering of skin, which forms for it a sort of protecting pouch,
and effectually prevents its being worn down by rubbing against the
ground. But when the Cat strikes its prey, it bends the paw upon the
Public-domain text, read in full here on John Shaqi.
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