From art, then, Darwin harks back once more to nature. He proceeds to
show that variability occurs among all wild plants and animals, though
not so frequently under ordinary circumstances as in the case of
domesticated species. Individual differences everywhere occur between
plant and plant, between animal and animal. Sometimes these differences
are so very numerous that it is impossible to divide the individuals at
all into well-marked kinds; for example, among British wild-roses,
brambles, hawkweeds and epilobes, with a few other very variable
families, Babington makes as many as 251 distinct species, where
Bentham gives only 112--a margin of 139 doubtful forms of shadowy
indefiniteness. Varieties, in fact, are always arising, and dominant
species in particular always tend to vary most in every direction. The
reason why variation is not so marked in the wild state as under
domestication is of course because the conditions are there less
diverse; but where the conditions of wild things are most diverse, as in
the case of dominant kinds, which range over a wide space of country or
of ocean, abundant individual variations habitually occur. Local
varieties thus produced are regarded by Darwin as incipient species:
they are the raw material on which natural selection gradually exerts
itself in the struggle for existence.
Granting individual variability, then, how do species arise in nature?
And how are all the exquisite adaptations of part to whole, and of whole
to environment, gradually initiated, improved, and perfected?
Here Malthus and the struggle for life come in to help us.
Public-domain text, read in full here on John Shaqi.
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