But towards the close of the eighteenth century, side by side with the general awakening of the human intellect and the arrival of a new era of free social investigation, which culminated in a fresh order of things, there was developed a more critical and sceptical attitude in the world of science, which soon produced a notable change of front among thinking naturalists as to the origin and meaning of specific distinctions. Buffon was the first great biological innovator who ventured, in very doubtful and tentative language, to suggest the possibility of the rise of species from one another by slow modification of ancestral forms. Essentially a popular essayist, writing in the volcanic priest-suppressed France of the _ancien régime_, during the inconsistent days of Louis XV. and Louis XVI., when it was uncertain whether novel and heterodox opinions would bring down upon their author fame and reputation or the Sorbonne and the Bastille, Buffon was careful to put his conjectural conclusions in a studiously guarded and often even ironical form. But time after time, in his great discursive work, the 'Histoire Naturelle' (published in successive volumes between 1749 and 1788), he recurs anew to the pregnant suggestion that plants and animals may not be bound by fixed and immovable limits of species, but may freely vary in every direction from a common centre, so that one kind may gradually and slowly be evolved by natural causes from the type of another. He points out that, underlying all external diversities of character and shape, fundamental likenesses of type occur in many animals, which irresistibly suggest the novel notion of common descent from a single ancestor. Thus regarded, he says, not only the ass and the horse (to take a particular passage) but even man himself, the monkeys, the quadrupeds, and all vertebrate animals, might be viewed as merely forming divergent branches of one and the same great family tree. Every such family, he believed, whether animal or vegetable, might have sprung originally from a single stock, which after many generations had here developed into a higher form, and there degenerated into a lower and less perfect type of organisation. Granting this--granting that nature could by slow variation produce one species in the course of direct descent from another unlike it (for example, the ass from the horse), then, Buffon observed, there was no further limit to be set to her powers in this respect, and we might reasonably conclude that from a single primordial being she has gradually been able in the course of time to develop the whole continuous gamut of existing animal and vegetable life. To be sure, Buffon always saves himself from censure by an obvious afterthought--'But no; it is certain from revelation that every species was directly created by a separate fiat.' This half-hearted and somewhat subrisive denial, however, must be taken merely as a concession to the Sorbonne and to the fashionable exegesis