Animal behavior; Animals -- Color; Plants -- Color
It is the enunciation and illustration of these two laws that form the
subject of the present treatise.
In the sequel we shall treat, in more or less detail, of each point as
it arises; but in order to render the argument clearer, this chapter is
devoted to a general sketch of my views.
Of the first great law but little need be said here, as it is almost
self-evident, and has never been disputed. It is true not only of the
upper and under-sides of animals, but also of the covered and uncovered
parts or organs.
For example, birds possess four kinds of feathers, of which one only,
the contour feathers, occur upon the surface and are exposed to the
light. It is in these alone that we find the tints and patterns that
render birds so strikingly beautiful, the underlying feathers being
invariably of a sober grey. Still further, many of the contour feathers
overlap, and the parts so overlapped, being removed from the light are
grey also, although the exposed part may be resplendent with the most
vivid metallic hues. A similar illustration can be found in most
butterflies and moths. The upper wing slightly overlaps the lower along
the lower margin, and although the entire surface of the upper wing is
covered with coloured scales, and the underwing apparently so as well,
it will be found that the thin unexposed margin is of an uniform grey,
and quite devoid of any pattern.
The law of structure, on the other hand, is an entirely new idea, and
demands more detailed explanation. Speaking in the broadest sense, and
confining ourselves to the animal kingdom, animals fall naturally into
two great sections, or sub-kingdoms, marked by the possession or absence
of an internal bony skeleton. Those which possess this structure are
known as _Vertebrata_, or backboned animals, because the
vertebral-column or backbone is always present. The other section is
called the _Invertebrata_, or backboneless animals.
Now, if we take the Vertebrata, we shall find that the system of
colouration, however modified, exhibits an unmistakably strong tendency
to assume a vertebral or axial character. Common observation confirms
this; and the dark stripes down the backs of horses, asses, cattle,
goats, etc., are familiar illustrations. The only great exception to
this law is in the case of birds, but here, again, the exception is more
apparent than real, as will be abundantly shown in the sequel. This
axial stripe is seen equally well in fishes and reptiles.
For our present purpose we may again divide the vertebrates into limbed
and limbless. Wherever we find limbless animals, such as snakes, the
dorsal stripe is prominent, and has a strong tendency to break up into
vertebra-like markings. In the limbed animals, on the other hand, we
find the limbs strongly marked by pattern, and thus, in the higher forms
the system of colouration becomes axial and appendicular.
Public-domain text, read in full here on John Shaqi.
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