Animal behavior; Animals -- Color; Plants -- Color
It may, however, be said, though I think not with justice, that we ought
not to take two species, however closely allied, to illustrate such a
point. But very good examples can be found in the same species. A common
German butterfly, _Araschnia Levana_, has two distinct varieties,
_Levana_ being the winter, and _prorsa_ the summer form; and between
these an intermediate form, _porima_, can be bred from the summer form
by keeping the pupæ cold. Dr. Weismann, who has largely experimented on
this insect, has given accurate illustrations of the varieties. Plate V.
is taken from specimens in our possession. In the males of both
_Levana_, Fig. 4, and _prorsa_, Fig. 1, the hind-wing has a distinct row
of spots, and a less distinct one inside it, and in the females of both
these are represented by dark stripes. In _porima_ we get every
intermediate form of spots and stripes, both in the male and female, and
as these were hatched from the same batch of eggs, or, are brothers and
sisters, it is quite impossible to doubt that here, at least, we have an
actual proof of the change of spots into stripes.
[Illustration: Fig. 1. Part of secondary feather of Argus Pheasant.
_a. a._ Elongated spots, incipient ocelli.
_b._ Interspaces.
_c. c._ Axial line.
_d. d._ Double spots, incipient ocelli.
_e._ Minute dottings.
_f. f._ Shaft.
_k. k._ Line of feathering.]
[Illustration: Fig. 2. Part of secondary wing feather of Argus
Pheasant.
_a._ Oval. Axis at right angles.
_b._ Round.
_c. c._ Shaft.
_d._ Imperfect ocellus.
_e._ Expansion of stripe.
_f._ Interspace.
_g._ Stalk.
_h._ Edge of feather.
_k._ Line of feathering.]
[Illustration: Plate V.
SEASONAL VARIETIES.]
The change of spots more or less irregular into eye-spots, or ocelli, is
equally clear; and Darwin's drawing of the wings of _Cyllo leda_[16]
illustrates the point well. "In some specimens," he remarks, "large
spaces on the upper surfaces of the wings are coloured black, and
include irregular white marks; and from this state a complete gradation
can be traced into a tolerably perfect ocellus, _and this results from
the contraction of the irregular blotches of colour_. In another series
of specimens a gradation can be followed from excessively minute white
dots, surrounded by a scarcely visible black line, into perfectly
symmetrical and larger ocelli." In the words we have put in italics
Darwin seems to admit these ocelli to be formed from blotches; and we
think those of the Argus pheasant can be equally shown to arise from
spots.
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