Comments on the Taxonomy and Geographic Distribution of North American MicrotinesHall, E. Raymond (Eugene Raymond)
Science
Comments on the Taxonomy and Geographic Distribution of North American Microtines
Hall, E. Raymond (Eugene Raymond)
Lemmings; Voles
The red-backed mouse of the Olympic Peninsula was originally accorded
specific rank. Currently it stands in the literature as a subspecies of
the wide-spread species _Clethrionomys gapperi_ because Dalquest (Univ.
Kansas Publ. Mus. Nat. Hist., 2:343, April 9, 1948) used the
name-combination _Clethrionomys gapperi nivarius_. Taylor and Shaw had
earlier (Occas. Papers Charles R. Conner Mus., 2:23, 1929) indicated
the same status by using the name _Evotomys gapperi nivarius_. Davis
(The Recent Mammals of Idaho, The Caxton Printers, Caldwell, Idaho, p.
306, April 5, 1939), however, indicated that the affinities of
_nivarius_ were with the _californicus_ [= _occidentalis_] group,
although he treated _nivarius_ as a distinct species. We have examined
two adult females (K. U. Nos. 10707 and 10708) of _nivarius_ from
Reflection Lake, 3800 ft., Jefferson County, Washington, and on the
basis of their thick, instead of thin, pterygoid processes concur with
Davis that the affinities of _nivarius_ are with the named kinds of
_Clethrionomys_ now arranged as subspecies of _Clethrionomys
occidentalis_, rather than with the kinds now arranged as subspecies of
_Clethrionomys gapperi_. Although we are aware that Dalquest (_op.
cit._:101-102) did not find actual intergradation between _nivarius_
and _Clethrionomys occidentalis occidentalis_--a ten-mile gap separated
their ranges--we prefer to use the name combination _Clethrionomys
occidentalis nivarius_. In doing so we recognize that intergradation
ultimately may be found between the two species _C. occidentalis_ and
_C. gapperi_; in that event the name _gapperi_ will apply as the name
of the species because it has priority over _occidentalis_.
The following named kinds of _Clethrionomys_ are considered to be
subspecies of _Clethrionomys occidentalis_:
CLETHRIONOMYS OCCIDENTALS OCCIDENTALIS (Merriam).
1890. _Evotomys occidentalis_ Merriam, N. Amer. Fauna, 4:25,
October 8, type from Aberdeen, Chehalis County, Washington.
1894. _Evotomys pygmaeus_ Rhoads, Proc. Acad. Nat. Sci.
Philadelphia, p. 284, October 23, type from mouth of Nisqually
River, Pierce County, Washington.
1929. _Evotomys gapperi occidentalis_, Taylor and Shaw, Occas.
Papers Charles R. Conner Mus., Washington State College, 2:23.
1948. _Clethrionomys californicus occidentalis_, Dalquest,
Univ. Kansas Publ., Mus. Nat. Hist., 2:344, April 9.
CLETHRIONOMYS OCCIDENTALIS CALIFORNICUS (Merriam).
1890. _Evotomys californicus_ Merriam, N. Amer. Fauna, 4:26,
October 8, type from Eureka, Humboldt County, California.
CLETHRIONOMYS OCCIDENTALIS CAURINUS (Bailey).
1898. _Evotomys caurinus_ Bailey, Proc. Biol. Soc. Washington,
12:21, January 27, type from Lund, east shore of Malaspina
Inlet, British Columbia.
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