Comments on the Taxonomy and Geographic Distribution of Some North American RodentsHall, E. Raymond (Eugene Raymond)
Science
Comments on the Taxonomy and Geographic Distribution of Some North American Rodents
Hall, E. Raymond (Eugene Raymond)
Mammals -- North America; Rodents
Harris (Occas. Papers Mus. Zool., Univ. Michigan, 266:1, June 28, 1933)
named _Sciurus variegatoides austini_ with type locality at Las Agujas,
Province of Puntarenas, Costa Rica. Later, in his revision of the
species _Sciurus variegatoides_, he (Misc. Publs. Mus. Zool., Univ.
Michigan, 38:19, September 7, 1937) referred specimens from Chomes,
Costa Rica, to _S. v. austini_ and (_op. cit._:24) specimens from
Puntarenas, Province of Puntarenas, to _S. v. rigidus_, an inland
subspecies. The geographic arrangement of these referred specimens
seemed to warrant a reconsideration of the material. We have examined
specimens of _S. variegatoides_ in the Museum of Zoology, University of
Michigan, from the following localities in Costa Rica: Puntarenas
(62703-62706), Las Agujas (65118 [type of _S. v. austini_],
59847-59850), Rio Las Agujas (65114-65117), Agua Caliente (66483),
Zarcero (75757-75761, 75765), Cartago (67546, 67547), and Esparta
(75762-75764). The specimens listed by Harris (_op. cit._, 1937:19) as
from Chomes, in the Museum of Zoology of the University of Michigan, are
not now in that museum and we have not seen them.
Harris (_op. cit._:19) characterized _S. v. austini_ as differing from
_S. v. rigidus_ in having brightly rufous legs (Ochraceous-Orange) in
_S. v. rigidus_ and a dorsal coloration resulting from a mixture of
shiny black and silver (Ochraceous-Orange mixed with black in _S. v.
rigidus_). We find that in the color of the legs of the paratypes of _S.
v. austini_ there is considerable variation ranging from bright rufous
in No. 65116 to much darker and duller in No. 59849. In six of the ten
specimens of the type series, the color is rufous, but in the other four
the color of the legs approaches and overlaps that found in the referred
specimens of _S. v. rigidus_. The color of the dorsum of _S. v. austini_
is also variable. No. 59850, for example, is dark brown and closely
resembles No. 75762, from Esparta, which was referred to _S. v.
rigidus_. Further, some specimens referred to _S. v. rigidus_ (67546 and
67547) have the bright-colored legs of _S. v. austini_ and some (75759,
for example) have the black-and-silver back of _austini_. We recognize
differences of an average sort between the now-available specimens of
the two alleged subspecies, but because of the individual variation that
exists, we feel that recognition of two subspecies is not indicated.
There is also some variation that is the result of wear and molt and one
of us (Kelson) feels that some of the differences are explainable on
this basis. Accordingly, we prefer to adopt a more conservative
taxonomic arrangement than that of Harris for this group of the Costa
Rican squirrels and arrange _Sciurus variegatoides austini_ Harris,
1933, as a synonym of _Sciurus variegatoides rigidus_ Peters, 1863.
~Thomomys bottae alienus~ Goldman
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