Contributions to the Theory of Natural Selection: A Series of EssaysWallace, Alfred Russel
Science
Contributions to the Theory of Natural Selection: A Series of Essays
Wallace, Alfred Russel
Natural selection
If we now pass to the next island (Celebes), separated from those last
mentioned by a strait not wider than that which divides them from each
other, we have a striking contrast; for with a total number of species
less than either Borneo or Java, no fewer than eighteen are absolutely
restricted to it. Further east, the large islands of Ceram and New
Guinea have only three species peculiar to each, and Timor has five. We
shall have to look, not to single islands, but to whole groups, in order
to obtain an amount of individuality comparable with that of Celebes.
For example, the extensive group comprising the large islands of Java,
Borneo, and Sumatra, with the peninsula of Malacca, possessing
altogether 48 species, has about 24, or just half, peculiar to it; the
numerous group of the Philippines possess 22 species, of which 17 are
peculiar; the seven chief islands of the Moluccas have 27, of which 12
are peculiar; and the whole of the Papuan Islands, with an equal number
of species, have 17 peculiar. Comparable with the most isolated of these
groups is Celebes, with its 24 species, of which the large proportion of
18 are peculiar. We see, therefore, that the opinion I have elsewhere
expressed, of the high degree of isolation and the remarkable
distinctive features of this interesting island, is fully borne out by
the examination of this conspicuous family of insects. A single
straggling island with a few small satellites, it is zoologically of
equal importance with extensive groups of islands many times as large
as itself; and standing in the very centre of the archipelago,
surrounded on every side with islets connecting it with the larger
groups, and which seem to afford the greatest facilities for the
migration and intercommunication of their respective productions, it yet
stands out conspicuous with a character of its own in every department
of nature, and presents peculiarities which are, I believe, without a
parallel in any similar locality on the globe.
Briefly to summarize these peculiarities, Celebes possesses three genera
of mammals (out of the very small number which inhabit it) which are of
singular and isolated forms, viz., Cynopithecus, a tailless Ape allied
to the Baboons; Anoa, a straight-horned Antelope of obscure affinities,
but quite unlike anything else in the whole archipelago or in India: and
Babirusa, an altogether abnormal wild Pig. With a rather limited bird
population, Celebes has an immense preponderance of species confined to
it, and has also six remarkable genera (Meropogon, Ceycopsis,
Streptocitta, Enodes, Scissirostrum, and Megacephalon) entirely
restricted to its narrow limits, as well as two others (Prioniturus and
Basilornis) which only range to a single island beyond it.
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