I should, be disposed to interpret the facts of the embryological
history and of the anatomy of the _Arthropoda_ in a different manner.
The _Copepoda_, the _Ostracoda_, and the _Branchiopoda_ are
the _Crustacea_ which have departed least from the embryonic or
_Nauplius_-forms; and, of these, I imagine that the _Copepoda_
represent the hypothetical _Archicarida_ most closely. _Apus_ and
_Sapphirina_ indicate the relations of these Archaeocarids with the
_Trilobita_, and the _Eurypterida_ connect the _Trilobita_ and the
_Copepoda_ with the _Xiphosura_. But the _Xiphosura_ have such close
morphological relations with the _Arachnida_, and especially with the
oldest known Arachnidan, _Scorpio_, that I cannot doubt the existence
of a genetic connection between the two groups. On the other hand, the
_Branchiopoda_ do, even at the present day, almost pass into the
true _Podophthalmia_, by _Nebalia_. By the _Trilobita_, again, the
_Archicarida_ are connected with such _Edriophthalmia_ as _Serolis_.
The _Stomapoda_ are extremely modified _Edriophthalmia_ of the
amphipod type. On the other side, the _Isopoda_ lead to the
_Myriapoda_, and the latter to the _Insecta_. Thus the Arthropod
phylum, which suggests itself to me, is that the branches of the
_Podophthalmia_, of the _Insecta_ (with the _Myriapoda_), and of the
_Arachnida_, spring separately and distinctly from the Archaeocarid
root--and that the _Zoaea_-forms occur only at the origin of the
Podophthalmous branch.
The phylum of the _Vertebrata_ is the most interesting of all, and is
admirably discussed by Professor Haeckel. I can note only a few points
which seem to me to be open to discussion. The _Monorhina_, having
been developed out of the _Leptocardia_, gave rise, according to
Professor Haeckel, to a shark-like form, which was the common stock
of all the _Amphirhina_. From this "Protamphirhine" were developed, in
divergent lines, the true Sharks, Rays, and _Chimaerae_; the Ganoids,
and the _Dipneusta_. The _Teleostei_ are modified _Ganoidei_. The
_Dipneusta_ gave rise to the _Amphibia_, which are the root of all
other _Vertebrata_, inasmuch as out of them were developed the first
_Vertebrata_ provided with an amnion, or the _Protamniota_. The
_Protamniota_ split up into two stems, one that of the _Mammalia_, the
other common to _Reptilia_ and _Aves_.
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