The only modification which it occurs to me to suggest in this
general view of the Phylogeny of the _Vertebrata_ is, that the
"Protamphirhine" was possibly more ganoid than shark-like. So far as
our present information goes the Ganoids are as old as the Sharks;
and it is very interesting to observe that the remains of the oldest
Ganoids, _Cephalaspis_ and _Pteraspis_, have as yet displayed no trace
of jaws. It is just possible that they may connect the _Monorhina_,
with the Sturgeons among the _Amphirhina_. On the other hand,
the Crossopterygian Ganoids exhibit the closest connection with
_Lepidosiren_, and thereby with the _Amphibia_. It should not be
forgotten that the development of the Lampreys exhibits curious points
of resemblance with that of the _Amphibia_, which are absent in
the Sharks and Rays. Of the development of the _Ganoidei_ we have
unfortunately no knowledge, but their brains and their reproductive
organs are more amphibian than are those of the Sharks.
On the whole, I am disposed to think that the direct stem of ascent
from the _Monorhina_ to the _Amphibia_ is formed by the Ganoids and
the Mudfishes; while the Osseous fishes and the Sharks are branches in
different directions from this stem.
What the _Protamniota_ were like, I do not suppose any one is in a
position to say, but I cannot think that the thoroughly Lacertian
_Protorosaurus_ had anything to do with them. The reptiles which are
most amphibian in their characters, and therefore, probably, most
nearly approach the _Protamniota_, are the _Ichthyosauria_ and the
_Chelonia_.
That the _Didelphia_ were developed out of some ornithodelphous form,
as Professor Haeckel supposes, seems to be unquestionable; but the
existing Opossums and Kangaroos are certainly extremely modified and
remote from their ancestors the "_Prodidelphia_," of which we have
not, at present the slightest knowledge. The mode of origin of the
_Monodelphia_ from these is a very difficult problem, for the most
part left open by Professor Haeckel. He considers the _Prosimiae_, or
Lemurs, to be the common stock of the _Deciduata_, and the _Cetacea_
(with which he includes the _Sirenia_) to be modified _Ungulata_. As
regards the latter question, I have little doubt that the _Sirenia_
connect the _Ungulata_ with the _Proboscidea_; and none, that the
_Cetacea_ are extremely modified _Carnivora_. The passage between the
Seals and the _Cetacea_ by _Zeuglodon_ is complete. I also think
that there is much to be said for the opinion, that the _Insectivora_
represent the common stock of the _Primates_ (which passed into
them by the _Prosimiae_), the _Cheiroptera_, the _Rodentia_, and the
_Carnivora_. And I am greatly disposed to look for the common root
of all the _Ungulata_, as well, in some ancient non-deciduate Mammals
which were more like _Insectivora_ than anything else. On the other
hand, the _Edentata_ appear to form a series by themselves.
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